Vol.:(0123456789) 1 3
Journal of Comparative Physiology B
https://doi.org/10.1007/s00360-018-1155-4
ORIGINAL PAPER
Iron storage disease (hemochromatosis) and hepcidin response to iron
load in two species of pteropodid fruit bats relative to the common
vampire bat
Iga M. Stasiak
1,2,4
· Dale A. Smith
1
· Tomas Ganz
3
· Graham J. Crawshaw
2
· Jutta D. Hammermueller
1
·
Dorothee Bienzle
1
· Brandon N. Lillie
1
Received: 27 October 2017 / Revised: 14 March 2018 / Accepted: 14 March 2018
© Springer-Verlag GmbH Germany, part of Springer Nature 2018
Abstract
Hepcidin is the key regulator of iron homeostasis in the body. Iron storage disease (hemochromatosis) is a frequent cause of
liver disease and mortality in captive Egyptian fruit bats (Rousettus aegyptiacus), but reasons underlying this condition are
unknown. Hereditary hemochromatosis in humans is due to defciency of hepcidin or resistance to the action of hepcidin.
Here, we investigated the role of hepcidin in iron metabolism in one species of pteropodid bat that is prone to iron storage
disease [Egyptian fruit bat (with and without hemochromatosis)], one species of pteropodid bat where iron storage disease
is rare [straw-colored fruit bat (Eidolon helvum)], and one species of bat with a natural diet very high in iron, in which iron
storage disease is not reported [common vampire bat (Desmodus rotundus)]. Iron challenge via intramuscular injection of
iron dextran resulted in signifcantly increased liver iron content and histologic iron scores in all three species, and increased
plasma iron in Egyptian fruit bats and straw-colored fruit bats. Hepcidin mRNA expression increased in response to iron
administration in healthy Egyptian fruit bats and common vampire bats, but not in straw-colored fruit bats or Egyptian fruit
bats with hemochromatosis. Hepcidin gene expression signifcantly correlated with liver iron content in Egyptian fruit bats
and common vampire bats, and with transferrin saturation and plasma ferritin concentration in Egyptian fruit bats. Induc-
tion of hepcidin gene expression in response to iron challenge is absent in straw-colored fruit bats and in Egyptian fruit bats
with hemochromatosis and, relative to common vampire bats and healthy humans, is low in Egyptain fruit bats without
hemochromatosis. Limited hepcidin response to iron challenge may contribute to the increased susceptibility of Egyptian
fruit bats to iron storage disease.
Keywords HAMP · Iron storage disease · Chiroptera · Iron metabolism · Hepcidin mRNA expression
Abbreviations
EFB Egyptian fruit bat
CVB Common vampire bat
SCFB Straw-colored fruit bat
Communicated by I. D. Hume.
Electronic supplementary material The online version of this
article (https://doi.org/10.1007/s00360-018-1155-4) contains
supplementary material, which is available to authorized users.
* Brandon N. Lillie
blillie@uoguelph.ca
Iga M. Stasiak
iga.stasiak@gov.sk.ca
Dale A. Smith
dalesmit@uoguelph.ca
Tomas Ganz
tganz@mednet.ucla.edu
Graham J. Crawshaw
crawshaw@bell.net
Jutta D. Hammermueller
jhammerm@uoguelph.ca
Dorothee Bienzle
dbienzle@uoguelph.ca
1
Department of Pathobiology, Ontario Veterinary
College, University of Guelph, 50 Stone Road, Guelph,
ON N1G 2W1, Canada
2
Toronto Zoo, 361A Old Finch Avenue, Toronto,
ON M1B 5K7, Canada
3
Department of Medicine, David Gefen School of Medicine
at UCLA, CHS 37-055, Los Angeles, CA 90095-1690, USA
4
Present Address: Saskatchewan Ministry of Environment,
Saskatoon, SK S7K 2H6, Canada