Journal of Pure and Applied Agriculture (2020) 5(1): 30-38
ISSN (Print) 2617-8672, ISSN (Online) 2617-8680
http://jpaa.aiou.edu.pk/
RESEARCH PAPER
Optimization of chitosan level to alleviate the drastic effects of heat stress
in cucumber (Cucumis sativus L.)
Mujahid Ali
1
*, Chaudhary Muhammad Ayyub
2
, Zahoor Hussain
1
, Rashid Hussain
2
and Shahla Rashid
1
1
Department of Horticulture, College of Agriculture, University of Sargodha, Sargodha, Pakistan
2
Institute of Horticultural Sciences, University of Agriculture, Faisalabad, Pakistan
*Corresponding author: Mujahid Ali (mujahidali2263@gmail.com)
Received: 21 February 2020; Accepted: 14 April 2020; Published online: 20 May 2020
Key Message: This study reveals the potential of chitosan
to alleviate the drastic effects of heat stress on cucumber
genotypes and optimization of chitosan for its effective and
economical use.
Abstract: Heat stress is a major concern during cucumber
production. To explore its production potential, chitosan
could play a vital role in alleviation of heat stress. This
study was planned to evaluate the potential of chitosan in
cucumber growth where it is more effective under high
temperature with two factorial under Completely
Randomized Design (CRD). Four genotypes were
previously screened out; two were tolerant (L3466, Desi-
cucumber) and two were sensitive (Suyo Long and
Poinsett). These genotypes were grown in the growth room
having a normal temperature (28
°
C/22
°
C day/night). One
day before heat treatment (40
º
C/32
º
C day/night), chitosan
foliar spray with different concentrations (0, 50, 100, 150,
200, 250, 300 ppm) was applied. It was found that 200 ppm of
chitosan level revealed significantly better results on growth
index (seedling length of shoot and root, the mass of fresh and
dry seedlings and the number of leaves per seedling) and
physiological index (chlorophyll contents and electrolyte
leakage). It was also found that heat sensitive genotype Suyo
Long gave the maximum seedling electrolyte leakage (74.03%)
followed by Poinsett (69.35%) at control (0 ppm chitosan). The
lowest seedling electrolyte leakage (28.85%) was noted in
Desi-cucumber followed by L3466 (30.16%) with foliar
application of 200 ppm of chitosan. So, chitosan spray at 200
ppm explored the maximum potential to alleviate the effect of
heat stress in cucumber genotypes. © 2020 Department of
Agricultural Sciences, AIOU
Keywords: Abiotic stress, Chitosan, Cucumber, Heat stress,
Optimization
To cite this article: Ali, M., Ayyub, C. M., Hussain, Z., Hussain, R., & Rashid, S. (2020). Optimization of chitosan level to
alleviate the drastic effects of heat stress in cucumber (Cucumis sativus L.). Journal of Pure and Applied Agriculture, 5(1), 30-
38.
Introduction
Cucumber (Cucumis sativus L.) has been playing a vital
role in food security. Its world production is 80.6 MT
(Food and Agricultural Organization Statistics
[FAOSTAT], 2016). Although, it is warm season crop yet
it is sensitive to heat stress (Zhang et al., 2012). Its annual
production is 71.7 MT globally (Khater, 2017). Cucumber
is native to subtropical and temperate zones and its best
growth and development is observed at 15-32
°
C.
However, high temperature above threshold level
deteriorates cucumber yield and quality (Zhao et al., 2011).
Characterization or screening of various genotypes of the
species against heat stress is inevitable at seedling level
(Shaheen et al., 2016; Sita et al., 2017) as heat stress is the
main hindrance in agriculture production (Schauberger et
al., 2017).
Agriculture and global warming are directly correlated.
The global temperature increased by 0.5
°
C in the last
hundred years. The global temperature would elevate by
1.8
°
C to 4
°
C during the next century (Intergovernmental
Panel on Climate Change [IPCC], 2014). The current
climatic model predicts that environmental temperature would
increase by 1.1-6.4
°
C if the CO
2
level doubles (Kim et al.,
2007). Global warming is becoming a sever issue (Haworth et
al., 2018). The high temperature is a principal cause of
unwanted alterations in the growth, development, and
physiology of plants (Shaked et al., 2004). The reproductive
phase is more sensitive as compared to the vegetative phase
because there are high losses at this stage (Meehl & Tebaldi,
2004). Heat stress proved to be the damaging effects during the
pollination process in chilies (Shaked et al., 2004), comparable
results were observed in bell pepper (Thuy & Kenji, 2015),
bean plant growth, development and yield (Omae et al., 2012),
and tomato yield and quality (Golam et al., 2012). When plants
suffer from heat stress, they enable the development of reactive
oxygen species that ultimately produce the oxidative stress
(Hasanuzzaman et al., 2012). It causes alterations in the
expression of genes that control heat tolerance potential
(Shinozaki & Yamaguchi-Shinozaki, 2007). In response, the
plants contest for the endurance of heat stress, such as
adjustment by changes in gene expression which becomes a
cause of heat tolerance to some extent (Moreno & Orellana,
2011). To avoid such situations in plants, foliar applications of