Received: 9 March 2021 Accepted: 18 August 2021
DOI: 10.1002/csc2.20628
Crop Science
ORIGINAL RESEARCH ARTICLE
Crop Breeding & Genetics
Multi-trait selection of bread wheat ideotypes for adaptation to
early sown condition
Md Farhad
1, 4
Shashi Bhushan Tripathi
1
Ravi Prakash Singh
2
Arun Kumar Joshi
3
Pradeep Kumar Bhati
3
Manish Kumar Vishwakarma
3
Suchismita Mondal
2
Anoop Anand Malik
1
Uttam Kumar
3
1
Dep. of Biotechnology, TERI School of
Advanced Studies, Vasant Kunj, New Delhi
110070, India
2
CIMMYT, Carretera México-Veracruz
Km. 45, El Batán, Texcoco, México C.P.
56237
3
Borlaug Institute for South Asia (BISA),
New Delhi 110012, India
4
Wheat Breeding Division, Bangladesh
Wheat and Maize Research Institute
(BWMRI), Dinajpur 5200, Bangladesh
Correspondence
Uttam Kumar, Borlaug Institute for South
Asia, New Delhi 11012, India.
Email: u.kumar@cgiar.org
Assigned to Associate Editor Paulo Teodoro.
Abstract
Genotypes developed for a particular mega-environment or management condition
may not suit different management or specific local environments. Planting earlier
than the recommended planting time is currently a new avenue to improve geno-
typic performance, supporting horizontal yield increases in the region when winter
is shortening. In India, wheat growers are looking at early planting because it can
use residual soil moisture from monsoons and escape terminal heat stress. A best
linear unbiased prediction (BLUP)-based multi-environmental stability analysis was
conducted on three sets of genotypes across three consecutive years (2017, 2018,
and 2019) under early and timely planting dates to identify genotypes for further
breeding. A number of traits were studied: phenological (early ground cover, days to
booting [DTB], days to heading, days to maturity [DAYSMT], boosting to heading
days [BTH], and grain filling duration [GFD]), plant stature (plant height, height up
to spike base, and spike length [SpkLng]), flag leaf (flag leaf length, flag leaf width,
and flag leaf area [FLGLFA]), and yield traits (thousand grain weight [TGW] and
grain yield). A significant genotypic effect was observed for all traits in the single
environment analysis. A genotype–environment interaction (GEI) was observed in
the mixed-effect model, except for FLGLFA in Season 2 and SpkLng in Season 3.
Residual components of variation were found to increase under early planting for all
studied traits due to exposure of genotypes to early heat and a prolonged growing
period. A higher GEI was observed in dissected phenological events such as BTH
and GFD. Among phenological traits, it was found that DTB, GFD, and DAYSMT
were strongly supporting selection gain throughout all seasons under early planting.
The genotypes with a more extended vegetative period and grain filling period tended
to have higher grain yield and TGW under early planting.
Abbreviations: BTH, booting to heading days; CIMMYT, International Maize and Wheat Improvement Center; DAYSMT, days to maturity; DTB, days to
booting; DTHD, days to heading; EGC, early ground cover; FA, factor analysis; FLGLFA, flag leaf area; GEI, genotype–environment interaction; GFD, grain
filling duration; GRYLD, grain yield; MGIDI, multi-trait genotype–idiotype distance index; PH, plant height; SpkLng, spike length; TGW, thousand grain
weight
© 2021 The Authors. Crop Science © 2021 Crop Science Society of America
Crop Science. 2021;1–16. wileyonlinelibrary.com/journal/csc2 1