Citation: Risoli, S.; Cotrozzi, L.;
Sarrocco, S.; Nuzzaci, M.; Pellegrini,
E.; Vitti, A. Trichoderma-Induced
Resistance to Botrytis cinerea in
Solanum Species: A Meta-Analysis.
Plants 2022, 11, 180. https://doi.org/
10.3390/plants11020180
Academic Editor: Paula Baptista
Received: 16 December 2021
Accepted: 4 January 2022
Published: 11 January 2022
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plants
Article
Trichoderma-Induced Resistance to Botrytis cinerea in Solanum
Species: A Meta-Analysis
Samuele Risoli
1,2
, Lorenzo Cotrozzi
2,3,
* , Sabrina Sarrocco
2,
* , Maria Nuzzaci
4
, Elisa Pellegrini
2,3
and Antonella Vitti
4,5
1
University School for Advanced Studies IUSS Pavia, Piazza della Vittoria 15, 27100 Pavia, Italy;
samuele.risoli@iusspavia.it
2
Department of Agriculture, Food and Environment, University of Pisa, Via del Borghetto 80, 56124 Pisa, Italy;
elisa.pellegrini@unipi.it
3
Nutrafood Research Center, University of Pisa, Via del Borghetto 80, 56124 Pisa, Italy
4
School of Agricultural, Forestry, Food and Environmental Sciences, University of Basilicata, Viale dell’Ateneo
Lucano 10, 85100 Potenza, Italy; maria.nuzzaci@unibas.it (M.N.); avitti@unisa.it (A.V.)
5
Department of Pharmacy, University of Salerno, Via Giovanni Paolo II, 132, 84084 Fisciano, Italy
* Correspondence: lorenzo.cotrozzi@unipi.it (L.C.); sabrina.sarrocco@unipi.it (S.S.);
Tel.: +39-050-221-0563 (L.C.); +39-050-221-6103 (S.S.)
Abstract: With the idea of summarizing the outcomes of studies focusing on the resistance induced
by Trichoderma spp. against Botrytis cinerea in tomato, the present paper shows, for the first time,
results of a meta-analysis performed on studies published from 2010 to 2021 concerning the cross-talk
occurring in the tomato–Trichoderma-B. cinerea system. Starting from an initial set of 40 papers, the
analysis was performed on 15 works and included nine parameters, as a result of a stringent selection
mainly based on the availability of more than one article including the same indicator. The resulting
work not only emphasizes the beneficial effects of Trichoderma in the control of grey mold in tomato
leaves (reduction in disease intensity, severity and incidence and modulation of resistance genes in
the host), but carefully drives the readers to reply to two questions: (i) What are the overall effects of
Trichoderma on B. cinerea infection in tomato? (ii) Do the main effects of Trichoderma differ based on
the tomato species, Trichoderma species, amount, type and duration of treatment? At the same time,
this meta-analysis highlights some weak points of the available literature and should be seen as an
invitation to improve future works to better the conceptualization and measure.
Keywords: grey mold; systemic resistance; pathogenesis-related (PR) genes; defense-signal transduc-
tion pathway; Trichoderma spp.
1. Introduction
Tomato (Solanum lycopersicum) is one of the most cultivated horticultural crops world-
wide, with more than 180 million tons produced over an area of about 5 million hectares in
2019 [1]. Tomato is also the most consumed vegetable worldwide and a major component
of the Mediterranean diet, mainly because of its remarkable nutraceutical properties due
to the considerable presence of antioxidant compounds, such as lycopene, ascorbic acid,
phenols, flavonoids and tocopherols [2]. At the same time, tomato is one of the most
economically important host plants of Botrytis cinerea (a pathogenic fungus belonging to
the phylum Ascomycota causing grey mold [3]).
Botrytis cinerea has been classified as the second most dangerous plant pathogen [4]
because of its wide host range (including more than 200 species over both temperate and
tropical areas) and its ability to attack leaves, stems, flowers and fruits, so determining
severe pre- and post-harvest losses [5,6], as well as the low efficacy of chemical control
due to its wide genetic variability and high capacity to acquire resistance against chemical
fungicides [7]. For these reasons, B. cinerea has received increasing attention, also becoming
Plants 2022, 11, 180. https://doi.org/10.3390/plants11020180 https://www.mdpi.com/journal/plants