a SciTechnol journal Research Article
Saumya and Yogesh Kumar, J Plant Physiol Pathol 2013, 1:3
http://dx.doi.org/10.4172/2329-955X.1000107
Journal of Plant
Physiology & Pathology
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Technology and Medicine
Arsenic Phytotoxicity in
Black Gram (Vigna mungo L.
Var. PU19) and its Possible
Amelioration by Phosphate
Application
Saumya Srivastava
1
* and Yogesh Kumar Sharma
1
Abstract
To study the toxic effect of arsenic on black gram (Vigna mungo
var. PU19) a pot experiment was conducted. Arsenic reduced the
germination percentage and root and shoot length of black gram.
Higher dose of arsenic (100 µM sodium arsenate) signifcantly
reduced the level of photosynthetic pigments. Increased oxidative
stress was observed as evident from enhanced level of lipid
peroxidation and electrolyte leakage percentage. However,
decreased catalase activity indicated its failure as antioxidant
enzyme against arsenic exposure. Treatment of phosphate
together with arsenic induced growth and metabolism in black
gram as compared to arsenic treatment alone.
Keywords
Arsenic; Vigna mungo; Pigment; Lipid peroxidation; Catalase;
Phosphate
Introduction
Arsenic, a group A carcinogen [1] is most commonly found in
oxidized states i.e arsenite and arsenate. Geologic and anthropogenic
activities like smelting operation, fossil fuel combustion [2] together
with use of arsenic based agrochemicals, fertilizer and disposal
of municipal & industrial wastes accounts for its widespread
contamination [3]. However, arsenic laden groundwater and soil are
considered major sources of contamination in the food and water
supplies. Also, irrigation of soils with arsenic laden water signifcantly
elevates the arsenic levels in the soil [4], rendering it unft for most
crop production, and severely afects the growth and development
of plants, and causes toxicity resulting in various physiological and
biochemical disorders [5]. Arsenic induced toxicity has been reported
in many plant species in the past [4,6-12].
Arsenic is quite similar to phosphate as both are placed in same
group, Va in the periodic table, have similar electron confguration,
chemical properties and compete for the same uptake carriers in
the root plasmalemma [13,14]. Tis also suggests its possible role in
ameliorating arsenic toxicity.
We selected black gram as its sensitivity/tolerance towards arsenic
*Corresponding author: Saumya Srivastava, Department of Botany, University of
Lucknow, Lucknow-226007, Uttar Pradesh, India; E-mail: sonata906@gmail.com
Received: February 25, 2013 Accepted: June 27, 2013 Published: July 01,
2013
toxicity is poorly understood. Black gram is an important pulse crop
in India and a rich source of protein. It is also grown for forage and
hay [15], and crop residues are an important feed for livestock. In
the present study, we investigated arsenic-induced toxicity in black
gram (Vigna mungo var. PU19) by studying the morphological and
physiological changes. Also, application of phosphate together with
arsenic was tested to determine the alterations it caused in various
growth and metabolic functions.
Methodology
Black gram (Vigna mungo var. PU19) seeds procured from
the Durga Egro Seeds Farm, Daliganj, Lucknow, Uttar Pradesh
were surface sterilized and presoaked in a petri-dish for 4 hours
containing sodium arsenate (Na
2
HASO
4
.7H
2
O) solution of following
concentrations as 0 µM(control), 50 µM and 100 µM with or without
40 ppm dipotassium hydrogen phosphate (K
2
HPO
4
) solution. Seeds
were then sowed in earthen pots having alluvial soil and compost (in
ratio 3:1). 500 ml of arsenic solution of above concentration was given
weekly in each pot kept in wire house. Plants were analyzed for all
parameters afer 45 days of sowing (except germination percentage
which was analyzed afer 7 days of sowing).
Germination percentage was calculated in each pot using formula-
Germination % = Number of germinated seeds X 100
Total number of seeds
Te root and shoot lengths of black gram plants (both treated
and control) were measured afer washing and rinsing plants with tap
and deionised distilled water respectively. Levels of photosynthetic
pigments (total chlorophyll, chlorophyll a and chlorophyll b) were
measured in black gram leaves according to Arnon [16]. 100 mg
leaves were crushed in 10 ml 80% chilled acetone. Te extracts were
centrifuged at 2000 g for 10 minutes. Absorbance of supernatant
was estimated spectrophotometrically at 645 nm and 663 nm using
a spectrophotometer (Toshniwal TSUV 75). Pigment contents
were expressed in terms of mg/g fresh weight of tissue. Carotenoid
contents were estimated according to the method of Duxbury and
Yentsch [17].
Electrolyte leakage percentage (ELP) was estimated according
to Sullivan & Ross [18]. Catalase activity was measured according
to the Euler, H. von & Josephson, K. method [19]. Catalase activity
was measured in terms of µM H
2
O
2
degraded/g fresh weight of tissue.
Peroxidase activity was estimated according to Luck, 1963 [20]. Lipid
peroxidation was evaluated to access the membrane damage in black
gram plants. For this, TBA (Tiobarbituric acid) test was used to
measure MDA (malondialdehyde) level which is an end product of
lipid peroxidation [21].
Te experiment was conducted in a completely randomized
design (CRD) with 3 replications. Te data were analyzed by One
Way ANOVA using sofware program Sigmastats 3.5. It was followed
by comparison of mean values using Holm Sidak method at p ≤ 0.05.
Results and Discussion
Germination percentage decreased with increasing arsenic