Introduction
The high incidence of motor side effects associated
with neuroleptic drug administration poses a signif-
icant problem for the treatment of schizophrenia.
These motor problems complicate treatment and are
often unresponsive to therapy.
1
There is still consid-
erable uncertainty about the mechanisms by which
neuroleptics produce these untoward side effects. It
has long been hypothesized that stimulation of super-
sensitive D2 receptors produces motor side effects
associated with neuroleptic use.
1
However, this and
other hypotheses invoking deficits in GABA, sero-
tonin, norepinephrine and acetylcholine transmission
as playing primary roles in this movement disorder
have provided inconclusive data regarding the mech-
anism(s) underlying the development of neuroleptic-
induced motor side effects.
1
One interesting idea that has emerged recently is
that effects of neuroleptics on the glutamate system
may underlie the development of motor side effects.
Chronic administration of typical neuroleptics has
effects on glutamate receptors that are not observed
with clozapine treatment.
2
Chronic administration
of haloperidol, but not clozapine, augments striatal
glutamate release.
3–5
It is thus possible that neuro-
leptic-induced enhancement of glutamate activity
contributes significantly to development of motor
side effects by causing prolonged low-level neuro-
toxicity at cortico-striatal synapses. This may occur
directly via neuroleptic-induced inhibition of gluta-
mate transport or indirectly via neuroleptic effects on
the dopamine system. Conventional neuroleptics
block D2 autoreceptors causing increased release of
dopamine.
6
Dopamine can, in turn, block glutamate
uptake,
7
thereby increasing synaptic concentrations
of glutamate and promoting excitotoxicity. Enhance-
ment of glutamatergic transmission could, with
either prolonged exposure or in individuals at risk
(i.e. with declining mitochondrial function due to
age) ultimately result in excitotoxicity. Enhancement
of glutamate activity in the presence of elevated
catecholamine metabolism could result in poten-
tiation of neurotoxicity.
8
Glutamate reuptake is the
primary mechanism for reducing synaptic concen-
trations of glutamate and preventing excitotoxicity.
9
Accordingly, the present study investigated anti-
psychotic drug effects on the gene expression of the
glutamate transporter GLT-1 in the striatum.
Materials and Methods
Adult male Sprague–Dawley rats (Taconic Farms)
weighing 300–350 g at the start of the study were
assigned to one of three treatment groups: halo-
peridol (0.5 mg/kg, i.p.) daily for 30 days, clozapine
(10.0 mg/kg, i.p.) daily for 30 days or saline (sterile,
i.p., in a volume similar to that used for drug admin-
Molecular Neuroscience
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© Rapid Science Publishers Vol 9 No 1 5 January 1998 133
RECENT reports have shown that typical neuroleptics
may enhance glutamatergic neurotransmission and that
these effects might in part underlie motor side effects of
chronic neuroleptic treatment. Since glutamate reuptake
is the primary mechanism for controlling extracellular
glutamate levels, the present study was conducted to
examine whether chronic neuroleptic exposure alters
gene expression for the glutamate transporter GLT-1 in
the striatum. Although both haloperidol and clozapine
treatment for 30 days significantly decreased GLT-1
expression from normal, the effects of haloperidol treat-
ment were consistently, and in the dorsal striatum,
significantly greater than those of clozapine. These find-
ings suggest that a deficiency in glutamate transport
may underlie the pathogenesis of neuroleptic-induced
movement disorders.
Key words: Clozapine; Glutamate; Haloperidol; Neuro-
leptics; Reuptake; Striatum
Chronic neuroleptic
treatment alters
expression of glial
glutamate transporter
GLT-1 mRNA in the
striatum
J. S. Schneider,
CA
Timothy Wade
and T. I. Lidsky
1
Department of Pathology, Anatomy and Cell
Biology, Thomas Jefferson University, 1020
Locust Street, Rm 521 JAH, Philadelphia, PA
19107;
1
Department of Psychobiology, Institute
for Basic Research, Staten Island, NY 10314,
USA
CA
Corresponding Author
Website publication 20 December 1997 NeuroReport 9, 133–136 (1998)