Plant and Soil 160: 57-66, 1994.
© 1994 KluwerAcademicPublishers.Printed in the Netherlands. PLSO 3195
Modification of soil aggregation by watering regime and roots growing
through beds of large aggregates
S.A. MATERECHERA l, J.M. KIRBY 2, A.M. ALSTON and A.R. DEXTER 3
Department of Soil Science, Waite Agricultural Research Institute, The University of Adelaide, Glen
Osmond, SA 5064, Australia. Present addresses: ~Crop Science Department, Bunda College of
Agriculture, University of Malawi, P.O. Box 219, Lilongwe, Malawi, eCSIRO Division of Soils, GPO
Box 639, Canberra, ACT 2601, Australia and 3Silsoe Research Institute, Wrest Park, Silsoe, Bedford
MK45 4HS, UK
Received 10 May 1993. Accepted in revised form 2l October 1993
Key words: aggregation, large aggregates, plant species, root growth, soil watering regimes, wetting
and drying
Abstract
The influence of root growth and soil watering regime on aggregation was studied under controlled
conditions. The study examined the influence of pea (Pisum sativum cv Greenfeast), ryegrass (Lolium
rigidum cv Wimmera) and wheat (Triticum aestivum cv Kite) roots on changes in aggregation and on the
properties of the aggregates. The soil was a non swelling red-brown earth which was either kept wet or
was allowed to wet and dry during the experiment. Root growth increased the percentage of small sized
aggregates (<18mm diameter), organic carbon, tensile strength and stability of aggregates in
comparison with a non planted soil. Changes in aggregate size distribution and properties of the
aggregates were related to root length density of the species and also to the soil watering regime. Root
length density was in the order ryegrass > pea > wheat. Wetting and drying of soil increased the strength
and stability of aggregates. Incubating aggregates allowed some roots to decompose but did not increase
the strength or stability of aggregates compared with unincubated soil. The results of this experiment
are of practical significance in soil structural management, and in studies of soil aggregation dynamics.
It may be possible to use plant roots to alter the size and properties of aggregates.
Introduction
Beds of sieved aggregates with a narrow dis-
tribution of sizes are often used in experiments
aimed at studying the dynamics of soil aggrega-
tion because they represent a reproducible and
well-defined structure (Braunack and Dexter,
1989). They also provide a means by which the
results of experiments conducted in different
places and times can be compared quantitatively.
Plant roots are known to grow preferentially in
weak aggregates or in voids between aggregates
(Dexter, 1986; McSweeney and Nausen, 1984).
When a plant root makes contact with the
surface of an individual aggregate in a bed of
aggregates larger than the diameter of the root,
it may penetrate, displace the aggregate or be
deflected by it (Misra et al., 1984; Whiteley and
Dexter, 1984). Penetration is determined by the
strength of the aggregate and by the angle of
incidence at which the root tip contacts the
aggregate (Dexter and Hewitt, 1978). The size of
the aggregate has also been reported to influence
the penetration of roots into aggregates (Logs-
don et al., 1987). The understanding of the
above processes has led to the development of