Journal of Oceanology and Limnology
https://doi.org/10.1007/s00343-019-7263-5
Metabolic and physiological regulation of Chlorella sp.
(Trebouxiophyceae, Chlorophyta) under nitrogen deprivation*
YONG Wai-Kuan
1, 2
, LIM Phaik-Eem
1, **
, VELLO Vejeysri
1, 3
, SIM Kae-Shin
3
,
ABDUL MAJID Nazia
3
, MUSTAFA Emienour Muzalina
4
, NIK SULAIMAN Nik Meriam
5
,
LIEW Kan-Ern
6
, CHEN Brenna Jia-Tian
7
, PHANG Siew-Moi
1, 3
1
Institute of Ocean and Earth Sciences, University of Malaya, 50603 Kuala Lumpur, Malaysia
2
Institute of Graduate Studies, University of Malaya, 50603 Kuala Lumpur, Malaysia
3
Institute of Biological Sciences, Faculty of Science, University of Malaya, 50603 Kuala Lumpur, Malaysia
4
School of Fisheries and Aquaculture Sciences, University of Malaysia, Terengganu, T145, 21300 Kuala Terengganu, Terengganu,
Malaysia
5
Department of Chemical Engineering, Faculty of Engineering, University of Malaya, 50603 Kuala Lumpur, Malaysia
6
Airbus Group Malaysia, Menara HLA, 3, Jalan Kia Peng, 50450 Kuala Lumpur, Malaysia
7
Aerospace Malaysia Innovation Centre, Level 1, MIGHT Building, 3517, Jalan Teknokrat 5, 63000 Cyberjaya, Selangor,
Malaysia
Received Sep. 11, 2017; accepted in principle Nov. 20, 2017; accepted for publication Mar. 2, 2018
© Chinese Society for Oceanology and Limnology, Science Press and Springer-Verlag GmbH Germany, part of Springer Nature 2018
Abstract A freshwater green microalgae Chlorella sp., UMACC344 was shown to produce high lipid
content and has the potential to be used as feedstock for biofuel production. In this study, photosynthetic
efficiency, biochemical profiles and non-targeted metabolic profiling were studied to compare between the
nitrogen-replete and deplete conditions. Slowed growth, change in photosynthetic pigments and lowered
photosynthetic efficiency were observed in response to nitrogen deprivation. Biochemical profiles of the
cultures showed an increased level of carbohydrate, lipids and total fatty acids, while the total soluble
protein content was lowered. A trend of fatty acid saturation was observed in the nitrogen-deplete culture
with an increase in the level of saturated fatty acids especially C16:0 and C18:0, accompanied by a decrease
in proportions of monounsaturated and polyunsaturated fatty acids. Fifty-nine metabolites, including
amino acids, lipids, phytochemical compounds, vitamins and cofactors were significantly dysregulated
and annotated in this study. Pathway mapping analysis revealed a rewiring of metabolic pathways in the
cells, particularly purine, carotenoid, nicotinate and nicotinamide, and amino acid metabolisms. Within
the treatment period of nitrogen deprivation, the key processes involved were reshuffling of nitrogen from
proteins and photosynthetic machinery, together with carbon repartitioning in carbohydrates and lipids.
Keyword: metabolic profiling; Chlorella sp.; nitrogen stress; lipid; fatty acid
1 INTRODUCTION
Microalgae are a potential feedstock for
triacylglycerol (TAG) and neutral lipids which are used
in biofuel production due to their fast growth rate, high
photosynthetic efficiency, high lipid yields and ability
to grow in a broad range of environmental conditions.
Various strategies, such as nutrient deprivation (e.g.
nitrogen starvation), temperature, salinity, pH, high
concentration of metal ion stress, and irradiation, have
been developed to improve the biomass yield and the
lipid content of microalgal cells (Sharma et al., 2012).
Nitrogen is an important building block for amino
acids, nucleic acids, enzymes and proteins. It is
assimilated into microalgal cells in the form of nitrate
or ammonium, and is involved in multiple metabolic
networks. While it is widely known that the deprivation
* Supported by the Aerospace Malaysia Innovation Centre & Airbus Group
Innovation (No. PV001-2013), the Ministry of Higher Education Malaysia
HICoE grant (No. IOES-2014H), the Fundamental Research Grant Scheme
(No. FP048-2016), and the University of Malaya UMCoE RU Grant (No.
RU009H-2015)
** Corresponding author: phaikeem@um.edu.my