Cancer Immunol. Immunother. 9, 245-253 (1980) ancer
mmunol_ogyand
mmunotherapy
© Springer-Verlag 1980
A Mouse Model for Immunotherapy of Osteosarcoma
III. Combined Therapy with Corynebacterium parvum and Prostaglandin Synthetase
Inhibitors
Paul Gatenby 1, Antony Basten 2, and Paul Creswick 2
1 Clinical Immunology, Royal Prince Alfred Hospital,
Camperdown, N.S.W. 2050
2 Immunology Unit, Medical School, Sydney University,
N.S.W. 2006, Australia
Summary. Various combinations of the prostaglan-
dins synthetase inhibitors, aspirin or indomethacin,
and Corynebacterium parvum were used as adjunctive
therapy to surgery in the treatment of the metastasizing
Dunn osteosarcoma C3H/HeJ mice. In doses corre-
sponding to those that could be tolerated by humans all
three agents given singly reduced the number and
estimated weight of metastases whether they were
administered pre-operatively or post-operatively.
When the two modalities of therapy were combined a
significant additive anti-tumour effect was observed
with both C. parvum and aspirin and C. parvum and
indomethacin, but only if treatment was commenced
pre-operatively. Similar results were obtained with two
other metastasizing turnouts, viz. the B16 melanoma
and the Lewis lung carcinoma. In the absence of any
evidence for an interaction between the prostaglandin
synthetase inhibitors and circulating tumour cells, it
was felt that the additive effect could be best explained
in terms of interference with prostaglandin-mediated
negative feedback of the anti-tumour action of C.
parvum.
Introduction
Prostaglandins (PGs) have recently been shown to
exert an influence on tumour growth and spread [22,
33, 36, 38, 39]. However, the mechanisms involved in
this interaction are complex since PGs, even of the
same series (PGEs) have been found to exert an
anti-tumour effect both in vivo [36] and in vitro [35],
while subverting anti-tumour responses in other
systems [22, 33]. The evidence for a tumour-poten-
tiating role of PGs comes largely from studies
showing that PG synthetase inhibitors, e.g., indo-
methacin or aspirin, can reduce the growth of
Reprint requests should be adressed to: P. Gatenby
tumours in certain animal systems [22, 26, 33]. The
mechanisms by which they do this is unclear, but it is
apparent that some tumours can secrete PGs [8, 39].
In addition, activated macrophages [19, 20, 30], T
cells [19], and even B cells [19] may produce PGs, and
there are ample studies in man and animals impli-
cating PGEs as local feedback inhibitors of T-cell
activation in vitro and in vivo [reviewed in 19]. The
general state of immunodepression that accompanies
disseminated malignancy has been shown to be
mediated by a PGE-dependent suppressor mecha-
nism in some animal [19, 33] and human turnouts
[20], but not in others [19]. Three potential sites exist
for the anti-turnout actions of PGEs. They may act
peripherally on mononuclear phagocytes [37] or
natural killer (NK) cells [7, 8] or may exert their
inhibitory effect centrally, by reducing the output of
mononuclear phagocytes from the myelomonocytic
progenitor cells in the bone marrow [24, 32]. Systemic
Corynebacterium parvum is thought to exert its
non-specific anti-tumour effect by increasing the
output of mononuclear phagocytes from the marrow
[6, 10, 12, 42] and/or by activating macrophages [3,
17], and NK cells via interferon [1, 2, 40]. These
mechanisms are illustrated schematically in Fig. 1,
together with the potential sites at which PGEs may
interfere with them. The synthesis of PGEs depends
on the enzyme prostaglandin synthetase or cycloox-
ygenase [28], and therefore any drugs which interfere
with the action of this enzyme will lead to a reduction
in PGEs. Consequently, an investigation of the
combined anti-tumour effect of C. parvum and
prostaglandin synthetase inhibitors seems logical on
theoretical grounds.
The PG synthetase inhibitors aspirin and indo-
methacin have in fact been shown to exert an
anti-tumour effect alone and to potentiate the
immunotherapeutic agents BCG and C. parvum in a
model involving flank tumours in C3H mice [26]. The
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