Plant Cell. Tissue and Organ Culture 33: 339-341, 1993.
© 1993 KluwerAcademic Publishers. Printedin the Netherlands.
In vitro propagation as an aid for cloning of Morus laevigata Wall
R. Islam, A. Zaman, O.I. Joarder & A.C. Barman
Department of Botany, University of Rajshahi, Rajshahi-6205, Bangladesh
Received 10 December 1991; accepted in revised form 7 December 1992
Key words: mulberry, micropropagation, nodal explants
Mulberry (Morus sp.) is the most important crop
in the sericulture industry because silkworms
(Bombyx mori L.) feed on its leaves. Since cross
pollination is the rule, enormous heterozygosity
occurs in this plant (Das 1991). Morus laevigata
is famous for quality timber and sweet edible
fruits about 6-12 cm long (Jain & Kumar 1989).
Again leaves of M. laevigata when fed to sil-
kworms produce a better cocoon than those fed
on M. alba (Mahmood et al. 1987). In spite of all
these good qualities, M. laevigata is very difficult
to propagate through cuttings because of its poor
rooting ability (Rajah & Ravindran 1989).
Clonal propagation of plants using tissue cul-
ture technique has many applications in ag-
ronomy (Murashige 1974), especially for woody
plants that are extremely difficult to propagate
by conventional means. Considerable work has
been done in the last few years on in vitro
propagation of M. nigra (Yadav et al. 1990), M.
indica (Patel et la. 1983) and M. alba (Hossain
et al. 1990). However, there is no report on in
vitro culture of M. laevigata. Here we describe
micropropagation using axillary buds from ma-
ture plants of M. laevigata to provide an alter-
native for its rapid clonal propagation.
Young shoots each having 4-5 nodes were
collected from a 12-year-old tree of M. laevigata
in June-July and thoroughly washed in running
tap water. Surface disinfestation was carried out
with 0.1% (w/v) HgC12 for 10 min after a brief
rinse in 70% (v/v) aq. ethanol. The material was
then washed with sterilized double-distilled
water for 10 rain giving 4-5 changes. Single node
explants were then excised and cultured on MS
medium (Murashige & Skoog 1962) containing
different concentrations (2.5-15 IxM) of cyto-
kinin, benzyladenine (BA) or kinetin individual-
ly or in combination with 0.5 ~xM of an auxin,
2,4-dichlorophenoxyacetic acid (2,4-D) or.
naphthaleneacetic acid (NAA). For adventitious
root induction, MS medium with half-strength
salts and sucrose with or without 0.5 ixM in-
dolebutyric acid (IBA) was used. The pH of the
medium was adjusted to 5.8 before autoclaving
and solidified with 7 g 1-1 Difco Bacto-agar. The
explants were cultured singly in 150 × 25mm
culture tubes each containing 15 ml of culture
medium and fitted with cotton plugs. The cul-
tures were grown at 25-+2°C at a relative
humidity of 55-65% (ambient conditions) under
a 16-h photoperiod with a light intensity of 50-
60~mol m 2 s-1 provided by warm white
fluorescent tubes. Data on days taken to bud
proliferation, percentage of proliferating cul-
tures, number of shoot per culture and shoot
length (cm) were recorded after 5 weeks of
culture. For each treatment 20 replicates were
used and each experiment was repeated at least
twice.
Establishment of explants in vitro from adult
trees was different due to excessive exudation of
a latex-like milky substances. Two to three
changes of media during the first week of culture
were found to be most beneficial for a satisfac-
tory response of explants in all instances. Nodal
explants cultured on different treatment combi-
nations showed their first response by enlarge-
ment and break of axillary buds. Rapid and early
proliferation was observed in media with lower
concentrations of cytokinins (Table 1). Absence
of growth regulators in the media delayed bud