pubs.acs.org/JAFC Published on Web 03/23/2010 © 2010 American Chemical Society
5374 J. Agric. Food Chem. 2010, 58, 5374–5379
DOI:10.1021/jf100413m
Bioaccessibility, Uptake, and Transport of Carotenoids from
Peppers (Capsicum Spp.) Using the Coupled in Vitro Digestion
and Human Intestinal Caco-2 Cell Model
LAURIE O’SULLIVAN,MARVIN A. JIWAN,TREVOR DALY,NORA M. O’BRIEN, AND
S. AISLING AHERNE*
School of Food and Nutritional Sciences, University College Cork, Cork, Republic of Ireland
Spanish bell peppers (Capsicum annuum L.) and chili peppers sourced from Kenya and Turkey
were analyzed for their carotenoid content, bioaccessibility, and bioavailability. The order of total
carotenoid content in peppers and their respective micelles was red > green > yellow. In terms of
cellular carotenoid transport as a percentage of original food and micelle content, the order was
yellow peppers > green > red; however, the opposite trend was seen for the actual amount of total
carotenoids transported by Caco-2 cells. Although lutein was generally the most abundant
carotenoid in the micelles (496.3-1565.7 μg 100 g
-1
), cellular uptake and transport of β-carotene
were the highest, 8.3-31.6 and 16.8-42.7%, respectively. Hence, the actual amount of carotenoids
present in the original food and respective micelles seems to reflect the amount transported by
Caco-2 cells. Therefore, color influenced the carotenoid profile, bioaccessibility, and bioavailability of
carotenoids rather than pepper type.
KEYWORDS: Bioaccessibility; bioavailability; Capsicum; Caco-2 cells; carotenoids; peppers
INTRODUCTION
Peppers (Capsicum annuum L.) are commonly consumed
vegetables because they provide a variety of color and flavor to
food products and meals. According to Sun et al. ( 1 ), the color of
sweet bell peppers is the major factor associated with consumer
purchasing decisions. Depending on flavor intensity and texture,
their culinary use changes from that as a vegetable (bell pepper)
to a spice (chili pepper) or a colorant (paprika). Peppers have
been reported to possess certain biological properties ( 1 , 2 ) that
may have positive effects on human health and chronic disease.
For instance, reduced risks of prostrate cancer have been asso-
ciated with increased intake of certain vegetables including bell
peppers ( 3 ). In addition, peppers have been shown to possess
radical scavenging activity ( 1 , 2 , 4 ) as well as prevent the oxidation
of cholesterol and docosahexaenoic acid (DHA) during heat-
ing ( 1 ). Peppers are one of the top 10 contributors to the dietary
intake of the carotenoids lutein and zeaxanthin in both the
Republic of Ireland and Spain ( 5 , 6 ). These plant foods are a
rich source of bioactive phytochemicals including capsaicinoids
and carotenoids ( 7 ).
Numerous epidemiological studies have shown an association
between the consumption of carotenoid-rich foods and a reduc-
tion in the risk of several human chronic diseases ( 8 , 9 ). The major
interest in studying carotenoids is because of their bioactive
effects, which include provitamin A activity, antioxidant actions,
immune modulation, and involvement in cell signaling ( 6 , 10 ).
Bioavailability is a critical feature in the assessment of the role
of dietary components, plant food consumption, and human
health ( 11 ). Therefore, to enhance our knowledge about the
amounts of carotenoids that are potentially available for absorp-
tion from commonly consumed carotenoid-rich foods such as
peppers, carotenoid bioavailability from these foods should be
investigated. In addition, factors that influence the content and
bioavailability of carotenoids from peppers should be identified
and established to evaluate the actual health effects obtained from
consuming such foods. Consequently, in the present study, we
determined the content, bioaccessibility, and bioavailability of
carotenoids from bell peppers (red, green, and yellow) and chili
peppers (red and green) that were available to the Irish consumer
at the time of testing, by using the in vitro digestion model
coupled with differentiated human intestinal Caco-2 cells.
Furthermore, we investigated if carotenoid content and bioacces-
sibility varied between chili peppers sourced from Kenya and
from Turkey.
MATERIALS AND METHODS
Material. All reagents including DMEM growth media, nonessential
amino acids, Hank’s balanced salts solution (HBSS), β-carotene, zeax-
anthin (Fluka), lutein (Fluka), and cholesterol esterase (Fluka) were
purchased from Sigma-Aldrich Chemical Co. (Dublin, Ireland). β-Cryp-
toxanthin (>95% purity) was purchased from LGC Prochem (Middlesex,
U.K.). Fetal bovine serum was sourced from Bio-Sciences (Co. Dublin,
Ireland). All solvents employed were of HPLC grade.
Sample Preparation and in Vitro Digestion. During one season,
uniformly ripe healthy peppers were purchased from a well-known super-
market chain. The red, green, and yellow bell peppers originated from
Spain. Chili peppers (red and green) were from Turkey and Kenya.
The peppers were not identical in genotype (i.e., seven different varieties).
*Corresponding author (telephone þ353 21 4902496; fax þ353 21
4270244; e-mail a.aherne@ucc.ie).