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Growth Study, Metabolite Development, and Organoleptic Profile
of a Malt-Based Substrate Fermented by Lactic Acid Bacteria
Lorenzo C. Peyer and Emanuele Zannini, School of Food and Nutritional Sciences, National University of Ireland,
University College Cork, College Road, Cork, Ireland; Fritz Jacob, Research Center Weihenstephan for Brewing and
Food Quality, Technische Universität München, Alte Akademie 3, 85354 Freising-Weihenstephan, Germany; and Elke K.
Arendt,
1
School of Food and Nutritional Sciences, National University of Ireland, University College Cork
ABSTRACT
J. Am. Soc. Brew. Chem. 73(4):303-313, 2015
The objective of this study was to investigate the fermentation of bar-
ley malt extract by four different lactic acid bacteria (LAB) species and to
analyze the effect on biological, physicochemical, and organoleptic attrib-
utes of the resulting beverages. Malt and wort were analyzed following
the Methodensammlung der Mitteleuropäischen Analysenkommission
(MEBAK). Sugars and organic acids were evaluated using HPLC, while
aroma compounds and ageing indicators were quantified using a head-
space GC-FID. Unhopped wort was able to support the growth of all LAB
during the whole experiment time (120 hr), with a cell count increase up
to 3.63 log orders. The strain-specific preferences for carbon and nitrogen
sources led to significant differences in the accumulation of metabolic
byproducts (i.e., organic acids and flavor-active compounds released in
the medium). LAB fermentation lowered the amount of some Strecker-
aldehydes (2- and 3-methylbutanal and 2-phenylacetaldehyde) carrying
“malty” off-flavors and increased dairy-related aroma compounds such as
diacetyl and acetoin (“buttery”). The fermented samples were sensorially
distinct and described with “bready”, “yeasty”, “yogurty”, or “honey”
attributes. In general, LAB fermentation conferred flavor complexity to
wort. Forced ageing of the fermented samples led to a large increase of
“staling” indicators but these samples improved organoleptic stability
compared with the unfermented control.
Keywords: Ageing, LAB, Lactic fermentation, Novel beverages, Sen-
sory, Strecker-aldehydes
During the last few decades, the accumulation of scientific evi-
dence linking nutrition and well-being has progressively steered
the choices of the modern consumer toward healthier and natural
food alternatives (26,68). This trend compels food companies to
continuously work on novel formulations or adapt older ones to
new standards (17). The increase in “functionality” is a wide-
spread and commercially attractive way to improve the potential
health benefits of food products (66). Functional beverages, in
particular, have recently been widely popularized in the form of
flavored waters, fruit juices, energy drinks, and teas (72).
Cereals carry beneficial functions in the form of dietary fiber
(1) and phytochemicals (e.g., sterols and phenolic acids) (2). There-
fore, they constitute an attractive substrate for the development of
novel, nonalcoholic, and functional beverages. The production of
cereal-based drinks is an interesting way to diversify the product
portfolio for many breweries and to relieve them from any major
economic losses due to the general decrease in beer consumption
(77). The implementation of this technology does not normally
require a radical change in the existing plant and can take ad-
vantage of any overcapacities present (43).
Because unprocessed cereal-based products are often consid-
ered bland or unpleasant in flavor (14), fermentation is often ap-
plied in order to improve the palatability (49,60). Lactic acid bac-
teria (LAB) can be used to increase the flavor complexity of these
substrates. This heterogeneous group of gram-positive LAB have
a long history of use in food products, and several of them have
been granted “generally recognized as safe” status by the Food
and Drug Administration. The use of LAB to ferment wort is ap-
plied to regulate the pH levels during the production of beer
aimed to be manufactured according to the German purity law
(47). LAB fermentation gives rise to a wide range of volatile and
nonvolatile compounds (e.g., organic acids, alcohols, aldehydes,
ketones, and carbonyl compounds) (54). Depending on the meta-
bolic features of the starter culture, as well as the cereal substrate
and the manufacturing processes involved, different strains of
LAB may generate characteristic and contrasting end products
(6). The knowledge of which organoleptic-active compounds are
being released during wort fermentation can help in the tailored
formulation of novel, specialty “sour beers” inspired by European
styles such as the Belgian Lambic or the German Berliner Weisse,
or from traditional fermented beverages (83). Malting of cereals
helps to provide better conditions for sustaining LAB fermenta-
tion due to the higher amount of assimilable nutrients (13) and
contributes to the aroma profile of the substrate (46).
Sensory evaluation plays an essential role in profiling the flavor
attributes and assessing the final consumer acceptance of a novel
product (11). Maintaining the smell and flavor quality from bot-
tling throughout the shelf-life of the product during storage repre-
sents an enormous challenge. Natural deterioration takes place
due to chemical reactions, mechanical stress during transporta-
tion, light influence, thermal stress caused by inadequate storage
conditions, and oxidative processes (57). Over the course of the
product shelf-life, cereal-based beverages commonly experience
an accumulation of staling compounds and a decrease in positive
organoleptic attributes (81). For these reasons, the understanding
of the evolution of the flavor profile during ageing is essential for
the commercial success of a product. Numerous studies and re-
views have been carried out on the origins of flavor deterioration
in beer (21,70,75,79) but only a few have been considered the
same problem for LAB-fermented cereal-based beverages (43).
The main aim of this work was to study the suitability of barley
malt-based wort to build a base for novel nonalcoholic beverages
fermented by LAB. Biological, physicochemical, and organolep-
tic changes were periodically studied for species of Weissella
cibaria, Lactobacillus brevis, L. reuteri, and L. plantarum and
related to the strain-specific metabolism. The flavor profile of the
fermented samples was compared before and after simulated age-
ing with the aim of understanding the main chemical transfor-
mations responsible for “staling”.
EXPERIMENTAL
Bacterial Cultures and Reagents
The microorganisms used as inoculum in this study (Table I) be-
long to the culture collection of the Cereal and Beverages Research
1
Corresponding author: Phone: +353 21 490 1449; Fax: +353 21 427 0213; E-mail:
e.arendt@ucc.ie
http://dx.doi.org/10.1094/ASBCJ-2015-0811-01
© 2015 American Society of Brewing Chemists, Inc.