~ 150 ~
Journal of Entomology and Zoology Studies 2018; 6(2): 150-156
E-ISSN: 2320-7078
P-ISSN: 2349-6800
JEZS 2018; 6(2): 150-156
© 2018 JEZS
Received: 01-01-2018
Accepted: 03-02-2018
Milagros M Greif
Biology Department, Cebu
Normal University, Cebu,
Philippines
Correspondence
Milagros M Greif
Biology Department, Cebu
Normal University, Cebu,
Philippines
Study on mosquito ovary using transmission
electron microscopy
Milagros M Greif
Abstract
The study focuses on the ovaries of mosquito using transmission electron microscopy. Mosquito that
were used comprised of six different strains, namely: Culex pipiens Complex, Aedes aegypti, Aedes
vexans, Ochlerotatus cantans, Ochlerotatus rusticus collected from the Philippines and Germany. A meal
of blood in the mosquito can alter a series of changes in the cell structure that leads to the formation of a
mature egg. After the mosquito has taken a blood meal, some prominent changes occur in the epithelial
follicles. The very prominent changes that occur are the development of large inter-cellular spaces and a
decrease in desmosomal connections. Several numbers of microvilli covered the area of the oocyte facing
the epithelial follicles of which during seven hour after blood meal, push towards the extra-cellular
spaces created by the separation of the epithelial follicle cells. The microvilli vary in length and are not
regularly distributed like those of the typical intestinal epithelium. A unit membrane is seen which later
fuses to form a yolk. A meal of blood or controlled diet leads to a series of changes in cell structures in
the reproductive organ of female mosquitoes that quickly results to the formation of matured eggs.
Another prominent changes observed in the ovary of mosquitoes having blood meal is the presence of
various un-oriented microvilli in the area of the oocyte which is adjacent to the epithelial follicles. These
microvilli can also be observed in ovary of mosquitoes having no blood meal but they are smaller in size.
Furthermore, pits or vesicular bodies are developed after a mosquito has taken a blood meal.
Keywords: mosquito, ovary, oogenesis, transmission electron microscopy
Introduction
Most of the insects including mosquitoes reproduce sexually. Adult mosquitoes, typically
obtain nutrients for survival and reproduction from three sources
[1, 2]
: (1) teneral reserves from
larval feeding on the microbiota and detritus in water, (2) nectar or other plant juices for
energy, and (3) blood taken only by females for egg production.
Mosquito ovaries are classified as meroistic type because they contain nurse cells as well as
oocytes. Furthermore, they are categorized as polytrophic because groups of nurse cells are
enclosed with an oocyte in each ovarian follicle. In contrast to other insects, like for instance
mayflies (Ephemeroptera) and cockroaches (Blattodea), no nurse cells are present within the
oocytes. This ovary is classified as telotrophic meroistic type. In these insects, the follicle cells
are equipped to carry out secretion and protein synthesis
[3]
.
A central chamber or calyx is present in each paired ovary. The calyx is lined by an epithelium
which contains muscle filaments that are also present in the lateral oviduct. Two tracheae enter
each ovary and branch into the tracheoles within the ovarioles. Each paired ovary consists of
variable numbers of tubular epithelial ovarioles where the oocytes are placed in a linear
sequence according to their stage of growth
[3]
.
The number of ovarioles ranges from 50 to 500 depending on the species, physiological stages
as well as the size of the individual female
[1]
. Each of the ovariole is enclosed by a sheath
which consists of cells forming a thin squamous mesothelium. Proximally, the ovarian sheath
forms the long suspensory ligament which inserts into the 4
th
abdominal tergite.
The ovariole contains a germarium as well as a maturing and presumptive follicle. In the
anterior part of each ovariole, are the differentiated cells of germarium through which the
mitotic divisions of germ cells takes place resulting in the formation of the oocytes. In
mosquitoes, these are accompanied by other cells which later perform the task of nursing or
supplying nutrients to the developing gametes. In other insects, however, special nurse cells
are absent and instead the nutritional responsiblity falls upon the epithelium of the oocyte
follicles
[3]
. In the posterior portion of the ovariole, approaching to the oviduct, lies the