Fish and Fisheries. 2018;1–13. wileyonlinelibrary.com/journal/faf
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1 © 2018 John Wiley & Sons Ltd
Received: 4 December 2017
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Revised: 11 March 2018
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Accepted: 8 May 2018
DOI: 10.1111/faf.12298
ORIGINAL ARTICLE
Larval abundance across the European eel spawning area: An
analysis of recent and historic data
Håkan Westerberg
1
| Michael J. Miller
2
| Klaus Wysujack
3
| Lasse Marohn
3
|
Marko Freese
3
| Jan-Dag Pohlmann
3
| Shun Watanabe
4
| Katsumi Tsukamoto
2
|
Reinhold Hanel
3
1
Department of Aquatic Resources, Institute
of Freshwater Research, Swedish University
of Agricultural Sciences, Drottningholm,
Sweden
2
Department of Marine Science and
Resources, College of Bioresource
Sciences, Nihon University, Fujisawa-shi,
Kanagawa, Japan
3
Thünen-Institute of Fisheries Ecology,
Hamburg, Germany
4
Department of Fisheries, Faculty of
Agriculture, Kindai University, Nara, Japan
Correspondence
Håkan Westerberg, Institute of Freshwater
Research, Stångholmsvägen 2, SE-178 93,
Drottningholm, Sweden.
Email: hakan.westerberg@slu.se
Funding information
German Federal Ministry of Food and
Agriculture; Ministry of Education, Culture,
Sports, Science and Technology of Japan,
Grant/Award Number: 26-252030
Abstract
The abundance and distribution of leptocephalus larvae of the European eel ( Anguilla
anguilla, Anguillidae) were examined using ten historic and recent Sargasso Sea expe-
ditions that were selected on the basis of having the largest number of sampling sta-
tions and highest catches. The surveys cover the period 1920–2014. Station data
were recalculated to the same unit of larval density per unit area, and the irregular
station positions were transformed to a regular spatial grid to allow calculation of
comparable measures of abundance of the youngest (0+) leptocephalus cohort. The
result is that the mean and maximum densities of 0+ leptocephali after 2007 on aver-
age have decreased by 70%–80% from the densities during the period before the
drastic decrease in glass eel recruitment, which started in the 1980s. This is of the
same magnitude as the change in spawning stock, if the total continental commercial
landings are used as a proxy. In the same period, the glass eel recruitment in Europe
has decreased by more than 95%. The conclusion is that a major cause for the recruit-
ment decrease may be an increased leptocephalus mortality during the oceanic phase
or a large geographic shift in glass eel arrival. Combining the survey data, the spatial
distribution of 0+ leptocephali was concentrated south of the northernmost front in
the Subtropical Convergence Zone, but high densities were also found far south of
the front in the western part of the distribution area and leptocephali were present
also north of the average frontal position.
KEYWORDS
Anguillid eels, larval distribution, leptocephalus, oceanic environment, population decline,
recruitment
1 | INTRODUCTION
The spawning area of the European eel, Anguilla anguilla, was discov-
ered about a century ago by the Danish scientist Johannes Schmidt
who revealed that the only place small larvae, called leptocephali,
could be collected was in the southern Sargasso Sea (Schmidt, 1923).
This was astonishing at the time and even today because it meant
that the reproductive stage silver eels must migrate many thousands
of kilometers to reach their distant oceanic spawning area (Righton
et al., 2012).
Schmidt’s sampling for leptocephali in the Sargasso Sea began
last century after no small leptocephali of A. anguilla were collected
in the Mediterranean Sea or offshore of western Europe (Schmidt,
1906, 1909, 1912a,b). His effort to find the spawning area of A. an-
guilla moved offshore in the North Atlantic Ocean after 40–60-mm
leptocephali of that species were collected in the central Atlantic