[CANCER RESEARCH 37, 460-463, February 1977]
SUMMARY
With a direct fluorescence assay, the levels of mixed
function oxidase activity were determined in mitogen-acti
vated human lymphocytes. The O-deethylation of ethoxyre
sorufin to resorufin was used to quantitate mixed-function
axidase activity. Ethoxyresarufin O-deethylase activity was
low to nondetectable in noninduced, mitogen-activated
cells, but it was readily detected in 3-methylcholanthrene,
mitogen-activated lymphocytes. The activity was: (a) de
pendent on assay time and number of lymphocytes; (b)
dependent on the presence of reduced nicotinamide ade
nine dinucleatide phosphate; (c) stable to freezing at —80°
for at beast 2 weeks; (d) reproducibly detected in duplicate
samples of blood from one individual when cultured and
assayed at the same time; but (e) quite variable in samples
of blood from one individual at different times. Since in
hepatic and pulmonary tissue of model animal systems
ethaxyresorufin is a specific substrate for cytochnome P
448-associated monooxygenases, the use of this chemical
could proffer an assay that specifically measures human
cytachnome P-448-assaciated activity.
INTRODUCTION
It has been suggested that a correlation exists (11) in man
between susceptibility to bronchogenic cancer (cigarette
smoke associated) and the inducibility by chemical cancina
gens of monooxygenase enzymes in cultured blood lym
phacytes that hydroxylate such carcinogens (2, 6, 8—1 1, 13,
26). This observation seems to be analogous to the situation
in model systems using inbred strains of mice in which the
activity of a specific monooxygenase, AHH3 is (a) host-gene
regulated (7, 17, 19, 20, 24, 25), (b) genetically linked to
cancer susceptibility caused by MC and benzo(a)pyrene (12,
15, 16, 18), and (c) specifically associated with the presence
of cytochrame P-448 (see review, Refs. 19 and 20).
Ethoxyresarufin can serve as a substrate far mixed-func
tian axidases in mat(3-5), mouse,4 or hamster (5)tissue, and
I This work was supported by National Cancer Institute Contracts N01-CP
33362, N01-CP-43309, and NO1-CP-55605and by contracts from the Council
for Tobacco Research. The mention herein of a proprietary product does not
constitute an endorsement by the United States Department of Agriculture.
2 Present address: Forensic Science Department, Karolinska Institutet,
S-10401 Stockholm, Sweden.
3 The abbreviations used are: AHH, aryl hydrocarbon hydroxylase; MC,
3-methylcholanthrene.
4 R. E. Kouri, L. M. Schechtman, T. Rude, C. E. McKinney, K. Nayar, and
M. D. Burke, manuscript in preparation.
Received January 19, 1976; accepted November 9, 1976.
its metabolism seems to be intimately associated with the
level of cytachrome P-448 (4, 5). Recent evidence4 suggests
that the metabolism of ethoxyresomufin in inbred strains of
mice is under the same genetic control as is the metabolism
of benza(a)pyrene via the AHH enzyme complex. Since the
assay for O-deethylation of ethaxymesomufin (3) is direct
(requiring no extraction steps), sensitive, and amenable to
automation and since ethaxyresarufin may be specifically
metabolized by only cytochrome P-448, this chemical may
serve as an effective alternate substrate [rather than
benza(a)pyrene] for determination of mixed-function oxi
dase activity in human tissue. This report describes such an
assay.
MATERIALS AND METHODS
Ethoxyresarufin and resarufin were prepared as previ
ously described (3). Lymphocytes were isolated from appar
ently healthy human volunteers (ages 24 to 39) who were
under no current drug regimen. All were nonsmokers ex
cept for Subject T. R. Lymphocytes were isolated in Ficoll
Hypaque gradients (specific gravity, 1.080) (Microbiological
Associates, Bethesda, Md.), washed once with Hanks' bal
anced salt solution, and cultured in 25-sq cm plastic flasks
in 8.0-mb aliquots at a density of 0.5 x 10@cells/mb. The
culture medium was Roswell Park Memorial Institute Me
dium 1640 supplemented with 17% fetal calf serum, 0.025 M
N-2-hydroxyethylpiperazine-N'-2-ethanesulfanic acid
buffer, 1% phytohemagglutinin M (Bumroughs-Wellcame,
Greenville, N. C.), 50 j.@gstreptomycin per ml, and 50 units
penicillin pen ml. Mitogen stimulation is a prerequisite far
the development of manoaxygenase activity in lymphocytes
(6, 26). After 72 hr of exposure to the mitagens, lymphocyte
cultures were treated with either 1.5 nmales MC pen ml
medium or an equivalent amount of the solvent acetone.
Acetone concentration was 0.2%. Cells were collected 24 hr
later and either assayed immediately amstoned as frozen
pellets in 0.2 ml 0.1 M Tnis-HCI, pH 8.5, at —80° for up to 4
weeks. Approximately 4 x 10@cells were frozen pentube.
For assay of frozen material , each cell pellet was thawed to
4°in the presence of 0.2 ml of 0.01 mM ethaxymesomufin
(dissolved in 0.1 M Tnis-HCI, pH 8.5), sonically disrupted
(three 2-sec bursts), and transferred to a 0.5- x 0.5-cm
microfluorametem cuvet. The fluorescence of the suspen
sian was measured at an emission wavelength of 586 nm
with an excitation wavelength of 520 nm, using an Aminco
Bowman spectrophatofluarometer with a sample compart
ment thermostatically regulated to 25°.After the fluanes
460
CANCER RESEARCH VOL. 37
3-Methylcholanthrene-'induced Monooxygenase (0-Deethylation)
Activity of Human Lymphocytes1
M. Danny Burke,2 Richard T. Mayer, and Richard E. Kourl
Biochemistry Department, University of Texas Health Science Center, Dallas, Texas 75235 fM. D. B.); Veterinary Toxicology and Entomology, Research
Laboratory, Agricultural Research Service, United States Department of Agriculture, College Station, Texas 77840 (R. T. M.J; and Department of Biochemical
Oncology, Microbiological Associates, Bethesda, Maryland 20014 (R. E. K.)
Research.
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