JOURNAL OF THE
WORLD AQUACULTURE SOCIETY
Vol. 41, No. 3
June, 2010
Virulence of Vibrios Isolated from Diseased Black Tiger Shrimp,
Penaeus monodon, Fabricius
Aseer Manilal, S. Sujith, Joseph Selvin
1
, Chippu Shakir, and R.
Gandhimathi
Department of Microbiology, Bharathidasan University, Tiruchirappalli, 620 024 India
G. Seghal Kiran
Department of Biotechnology, Bharathidasan University, Tiruchirappalli, 620 024 India
Abstract
Vibrios are prominently opportunistic, but they may act as primary and secondary pathogenic
invaders of shrimps in farms worldwide. However, little is known about the virulence of diverse
vibrios harbored in shrimp farms. The isolation of secondary/opportunistic Vibrio pathogens was
carried out from the shrimp farms located in peninsular India, reported to have mass mortalities
because of White-spot syndrome virus and shell disease outbreaks. The isolates were identified and
grouped into six species including luminescent Vibrio harveyi, Vibrio alginolyticus, Vibrio vulnificus,
Vibrio fischeri, Vibrio parahaemolyticus, and Photobacterium damsela. To make a detailed study on
their virulence, the isolates were screened in vitro for the production of extracellular factors. In the
present study, 68.9% of the shrimp Vibrio isolates were found to be positive for both phospholipase
and hemolysin production. The results showed that various factors including hydrophobicity, presence
of capsule, and production of extracellular enzymes such as lipase, protease, phospholipase, and
hemolysin were responsible for pathogenesis. Using the bacterial adherence to hydrocarbons, most
of the strains were classified as highly hydrophobic. In vivo challenge experiments using the Vibrio
species and the extracellular products demonstrated the pathogenicity of vibrios. In the present study,
we established the shrimp farm water isolates V. vulnificus, V. fischeri, and P. damsela as moderately
opportunistic shrimp pathogens.
Members of the genus Vibrio are autochtho-
nous bacterial flora in the aquatic ecosystem,
and quite a few of them are associated with
infections in humans and aquatic animals. They
are the normal bacterial flora of shrimp and
the culture environment (Jiravanichpaisal et al.
1994; Otta et al. 1999), but they often act as
secondary or opportunistic pathogens that cause
mortality ranging from few to 100% in affected
populations under stress (Lightner 1988). Vib-
riosis has been implicated as the cause of major
mortality in juvenile penaeid shrimp (Light-
ner and Redman 1994). Vibrio spp. are most
often considered opportunistic pathogens in
shrimp, but primary disease caused by highly
virulent strains has also been reported (De
1
Corresponding author.
La Pe˜ na et al. 1992; Ishimaru et al. 1995).
Reports on the epizootic luminescent bacte-
rial diseases, especially in the shrimp farms
of Asian countries, have been connected to a
decrease in the shrimp production and inten-
sive rearing systems (Karunasagar et al. 1994).
Luminescent vibriosis is mainly caused by Vib-
rio harveyi and Vibrio campbellii and occa-
sionally by Vibrio splendidus, which can infect
larval juveniles and adult stages of penaeid
shrimp (Gomez-gill et al. 1998; Lavilla-pitogo
et al. 1998). In India, luminous V. harveyi
and Vibrio alginolyticus were reported as pre-
dominantly opportunistic and secondary shrimp
pathogens of highly devastating mid-culture
outbreaks (Selvin and Lipton 2003a; Selvin
et al. 2005). In this report, the disease caused
by vibrios because of deteriorated water quality,
© Copyright by the World Aquaculture Society 2010
332