Andrologia. 2019;00:e13393. wileyonlinelibrary.com/journal/and
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https://doi.org/10.1111/and.13393
© 2019 Blackwell Verlag GmbH
1 | INTRODUCTION
Semen cryopreservation and artificial insemination can be used to
control spread of many infectious diseases and promote dissemi‐
nation of superior genetics (Dalal, Ajeet, Ravi, Gyan, & Chandolia,
2018). However, utilisation of this assisted reproductive technology
depends on quality of frozen‐thawed semen, which can be influ‐
enced by many factors, including sperm cryoresistance (Inanc, Tekin,
et al., 2018). For example, changes in lipid composition of sperm
membranes affect quality of frozen‐thawed spermatozoa (Amorim,
Graham, Spizziri, Meyers, & Torres, 2009).
Novel freezing methods and semen extenders with various
supplements are still being investigated to further improve quality
of cryopreserved ram spermatozoa (Blackburn, 2004). Adding an‐
tioxidants to extenders for chilling or freezing semen may confer
antioxidant protection and improve sperm motility, mitochondrial
activity and viability (Bucak, Coyan, Ozturk, Gungor, & Omur,
2012). Lipid composition (lipid/cholesterol ratio) also affects abil‐
ity of spermatozoa to capacitate and undergo an acrosome reac‐
tion (He, Bailey, & Buhr, 2001; Moore, Squires, & Graham, 2005).
Specifically, ram spermatozoa are sensitive to cryopreservation‐
induced damage, due to their low phospholipid/cholesterol ratio
(Inanc, Uysal, & Ata, 2018). Furthermore, cryopreserved sperma‐
tozoa are more sensitive to oxidation than fresh spermatozoa, due
to less effective intracellular antioxidant defence mechanisms
(Chandra et al., 2012).
Received: 8 May 2019
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Revised: 18 July 2019
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Accepted: 19 July 2019
DOI: 10.1111/and.13393
ORIGINAL ARTICLE
Gallic and carnosic acids improve quality of frozen‐thawed ram
spermatozoa
Sukru Gungor
1
| Muhammed E. Inanc
1
| Caner Ozturk
2
| Firat Korkmaz
3
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Ilktan Bastan
3
| Beste Cil
4
| John P. Kastelic
5
1
Department of Reproduction and Artificial
Insemination, Faculty of Veterinary
Medicine, Burdur Mehmet Akif Ersoy
University, Burdur, Turkey
2
Department of Reproduction and Artificial
Insemination, Faculty of Veterinary
Medicine, Aksaray University, Aksaray,
Turkey
3
International Center for Livestock Research
and Training, Ankara, Turkey
4
Department of Reproduction and Artificial
Insemination, Faculty of Veterinary
Medicine, Ankara University, Ankara, Turkey
5
Faculty of Veterinary Medicine, University
of Calgary, Calgary, AB, Canada
Correspondence
Sukru Gungor, Department of Reproduction
and Artificial Insemination, Faculty of
Veterinary Medicine, Burdur Mehmet Akif
Ersoy University, 15030, Burdur, Turkey.
Email: sukrugungor85@gmail.com
Abstract
The objective was to determine effects of gallic acid (GA) and carnosic acid (CA),
present in carob pods and rosemary extract respectively, on frozen‐thawed ram sper‐
matozoa. Thirty ejaculates were collected from five Merino rams, pooled, diluted in
Tris‐based extender and divided into five equal portions containing: 0.05 or 2 mM of
GA; 0.05 or 0.2 mM of CA; or no additive (control). Extended semen was equilibrated
at +4°C, loaded into straws, held 5 cm above liquid nitrogen for 12 min then plunged.
Computer‐aided sperm analysis was used to assess motility, whereas flow cytometry
was used to assess high mitochondrial membrane potential (HMMP) and percentages
of spermatozoa with plasma membrane and acrosome integrity (PMAI). Spermatozoa
supplemented with 2 mM GA had greater total motility than control spermatozoa
(39.9 ± 3.01 vs. 29.2 ± 1.31%, mean ± SEM, p < .05). The PMAI was greatest in 0.2 mM
CA (13.3 ± 0.68%), whereas HMMP was highest in 0.05 mM CA but lowest in control
(22.9 ± 4.95 and 11.4 ± 3.64% respectively; p < .05). In conclusion, for cryopreserva‐
tion of ram semen in Tris‐based extender, supplementation with 2 mM GA increased
post‐thaw motility, whereas supplementation with 0.05 mM CA enhanced mitochon‐
drial function.
KEYWORDS
carnosic acid, computer‐assisted sperm analysis, flow cytometry, gallic acid