Vol.:(0123456789) 1 3
Journal of Molecular Histology
https://doi.org/10.1007/s10735-020-09888-3
ORIGINAL PAPER
Integrin‑linked kinase (ILK) regulates KRAS, IPP complex and Ras
suppressor‑1 (RSU1) promoting lung adenocarcinoma progression
and poor survival
Sofa Nikou
1
· Marina Arbi
2
· Foteinos‑Ioannis D. Dimitrakopoulos
3
· Chaido Sirinian
3
· Panagiota Chadla
1
·
Ioanna Pappa
1
· Giannoula Ntaliarda
4
· Georgios T. Stathopoulos
4,5
· Helen Papadaki
1
· Vasiliki Zolota
6
·
Zoi Lygerou
2
· Haralabos P. Kalofonos
3,7
· Vasiliki Bravou
1
Received: 26 April 2020 / Accepted: 13 June 2020
© Springer Nature B.V. 2020
Abstract
Integrin-linked kinase (ILK) forms a heterotrimeric protein complex with PINCH and PARVIN (IPP) in Focal Adhesions
(FAs) that acts as a signaling platform between the cell and its microenvironment regulating important cancer-related func-
tions. We aimed to elucidate the role of ILK in lung adenocarcinoma (LUADC) focusing on a possible link with KRAS
oncogene. We used immunohistochemistry on human tissue samples and KRAS-driven LUADC in mice, analysis of large
scale publicly available RNA sequencing data, ILK overexpression and pharmacological inhibition as well as knockdown of
KRAS in lung cancer cells. ILK, PINCH1 and PARVB (IPP) proteins are overexpressed in human LUADC and KRAS-driven
LUADC in mice representing poor prognostic indicators. Genes implicated in ILK signaling are signifcantly enriched in
KRAS-driven LUADC. Silencing of KRAS, as well as, overexpression and pharmacological inhibition of ILK in lung cancer
cells provide evidence of a two-way association between ILK and KRAS. Upregulation of PINCH, PARVB and Ras sup-
pressor-1 (RSU1) expression was demonstrated in ILK overexpressing lung cancer cells in addition to a signifcant positive
correlation between these factors in tissue samples, while KRAS silencing downregulates IPP and RSU1. Pharmacological
inhibition of ILK in KRAS mutant lung cancer cells suppresses cell growth, migration, EMT and increases sensitivity to
platinum-based chemotherapy. ILK promotes an aggressive lung cancer phenotype with prognostic and therapeutic value
through functions that involve KRAS, IPP complex and RSU1, rendering ILK a promising biomarker and therapeutic target
in lung adenocarcinoma.
Keywords ILK · KRAS · Lung cancer · PARVB · PINCH · Ras suppressor-1
Abbreviations
ECM Extracellular matrix
EMT Epithelial to mesenchymal transition
ERK1/2 Extracellular regulated kinase
Electronic supplementary material The online version of this
article (https://doi.org/10.1007/s10735-020-09888-3) contains
supplementary material, which is available to authorized users.
* Vasiliki Bravou
vibra@upatras.gr
1
Department of Anatomy-Histology-Embryology, Medical
School, University of Patras, 26500 Patras, Greece
2
Department of General Biology, Medical School, University
of Patras, 26504 Patras, Greece
3
Clinical and Molecular Oncology Laboratory, Division
of Oncology, Medical School, University of Patras,
26504 Rio, Greece
4
Laboratory for Molecular Respiratory Carcinogenesis,
Department of Physiology, Faculty of Medicine, University
of Patras, 2504 Rio, Achaia, Greece
5
Comprehensive Pneumology Center (CPC) and Institute
for Lung Biology and Disease (iLBD), University Hospital,
Ludwig-Maximilians University and Helmholtz Zentrum
München, Member of the German Center for Lung Research
(DZL), Munich, Bavaria, Germany
6
Department of Pathology, University Hospital of Patras,
26504 Patras, Greece
7
Division of Oncology, Department of Internal Medicine,
University Hospital of Patras, 26504 Rio, Greece