RESEARCH
PAPER
Role of environmental, historical and
spatial processes in the structure of
Neotropical primate communities:
contrasting taxonomic and phylogenetic
perspectives
Maria Mercedes Gavilanez* and Richard D. Stevens
Department of Biological Sciences, Louisiana
State University, Baton Rouge, LA, USA
ABSTRACT
Aim To evaluate relative importance of niche, historical and spatial processes on
the taxonomic and phylogenetic structure of primate communities.
Location Neotropics.
Methods Data on community composition for 74 sites were gathered from the
literature. Communities were characterized based on taxonomic and phylogenetic
composition. Three predictive matrices were used as explanatory variables repre-
senting ecological (environmental), historical (riverine barriers and Pleistocene
refugia) and dispersal-based spatial hypotheses (spatial principal coordinates of
neighbour matrices vectors based on geographic coordinates). Variation partition-
ing analyses were used to decouple independent and shared effects. Permutation
procedures based on redundancy analysis were used to determine if explained
variation was statistically significant.
Results Forty-nine per cent of variation in taxonomic structure and 66% of
variation in phylogenetic structure was explained by selected predictor variables.
Independent effects of spatial variables explained the largest portion of variation in
both diversity metrics. Fractions representing shared effects of historical and spatial
variables, and of all variables combined, were also large and significant. Independ-
ent effects of environmental and historical variables were small to non-existent
(< 4%). When analysing each phylogenetic metric independently the relative con-
tributions of shared fractions changed, although the independent spatial fraction
remained the most important predictor.
Main conclusions Our results show that purely spatial processes, such as disper-
sal limitation, may play a stronger role in structuring primate communities than
niche mechanisms and historical events. Furthermore, we find that the influence of
ecological and evolutionary mechanisms is conflated with spatial processes, sug-
gesting that community structure is determined jointly by spatial mechanisms
reflecting environmental gradients and biogeographical processes. Although the
relative contributions of each predictor variable were similar between taxonomic
and phylogenetic metrics, a deeper examination of phylogenetic metrics suggests
that ecological, historical and spatial mechanisms interact in complex ways to
determine current patterns of phylogenetic community structure.
Keywords
Community structure, dispersal-based processes, species–environment
relationships, historical processes, Neotropical primates, phylogenetic diversity.
*Correspondence: Maria Mercedes Gavilanez,
Department of Biological Sciences, Louisiana
State University, 107 Life Sciences Building,
Baton Rouge, LA 70803, USA.
E-mail: mgavil2@tigers.lsu.edu
Global Ecology and Biogeography, (Global Ecol. Biogeogr.) (2012) ••, ••–••
© 2012 Blackwell Publishing Ltd DOI: 10.1111/geb.12011
http://wileyonlinelibrary.com/journal/geb 1