Citation: Didi, D.A.; Su, S.; Sam, F.E.;
Tiika, R.J.; Zhang, X. Effect of Plant
Growth Regulators on Osmotic
Regulatory Substances and
Antioxidant Enzyme Activity of
Nitraria tangutorum. Plants 2022, 11,
2559. https://doi.org/10.3390/
plants11192559
Academic Editors: Massimiliano
Tattini, Antonella Gori and Luana
Beatriz dos Santos Nascimento
Received: 5 September 2022
Accepted: 24 September 2022
Published: 28 September 2022
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plants
Article
Effect of Plant Growth Regulators on Osmotic Regulatory
Substances and Antioxidant Enzyme Activity of
Nitraria tangutorum
Dom Alizet Didi
1
, Shiping Su
1,
*, Faisal Eudes Sam
2
, Richard John Tiika
1
and Xu Zhang
1
1
College of Forestry, Gansu Agricultural University, Lanzhou 730070, China
2
College of Enology, Northwest A&F University, Yangling 712100, China
* Correspondence: susp008@163.com
Abstract: Plant growth regulators (PGRs) are natural hormones and synthetic hormone analogues.
At low concentrations, PGRs have the ability to influence cell division, cell expansion, and cell
structure and function, in addition to mediating environmental stress. In this study, experiments
were conducted to determine how exogenous PGRs indole acetic acid (IAA), abscisic acid (ABA), and
gibberellic acid (GA) influenced osmotic regulatory substances and activity of antioxidant enzymes
in Nitraria tangutorum. Using a completely randomized design, IAA, ABA, and GA3 were applied
as foliar spray at concentrations of 50 mg/L, 100 mg/L, 150 mg/L, and 200 mg/L to N. tangutorum
shrubs. Some selected shrubs did not receive any treatment and served as the control (Ck). The results
showed that the foliar spray of IAA, ABA, and GA3 significantly increased the content of osmotic
regulatory substances (soluble sugar, soluble protein, and proline) and antioxidant enzymes (SOD
and POD) at most concentrations. In addition, the malondialdehyde (MDA) content significantly
reduced after treatment, but after regrowth of coppiced shrubs, lipid peroxidation increased and
was still lower than Ck. Our study provides evidence that 100 mg/L 150 mg/L, and 200 mg/L
concentrations of IAA, ABA, and GA3 treatments are effective for enhancing osmotic regulatory
substances and the activity of antioxidant enzymes in N. tangutorum, which offers an effective strategy
not only for increasing tolerance to abiotic and biotic stresses, but also improving the adaptability of
N. tangutorum shrubs to the environment.
Keywords: plant growth regulator; abscisic acid; indole acetic acid; gibberellic acid; osmotic
regulatory substances; antioxidant enzymes; Nitraria tangutorum Bobr.
1. Introduction
Plant growth regulators (PGRs) were first discovered in plants in the 20th century.
PGRs are a variety of diverse molecules known as plant hormones that act as chemical
messengers to control plant growth and development [1,2]. PGRs exist in both natural
(extracted) and synthetic (manufactured) forms. Although a synthetically produced plant
growth regulator is structurally identical to a plant regulator or hormone, it is not consid-
ered to be one [2]. The production and growth of roots, shoots, buds, flowers, and fruits can
be controlled by the use of synthetic PGRs [3]. PGRs are compounds that promote plant
growth and increase their stress tolerance under various stress conditions [4,5]. PGRs affect
a variety of plant traits, including plant height, number of leaves, leaf area index, dry matter,
chlorophyll concentration, and photosynthetic parameters, among others [6]. PGRs can
also improve plant tolerance to various abiotic stimuli, increase antioxidant capacity, and
accelerate plant growth [7]. PGRs have been synthesized and can be exogenously applied to
plants to control their growth and development. PGRs come in a variety of forms, including
auxins such as indoleacetic acid (IAA), abscisic acid (ABA), and gibberellic acid (GA).
In plants, IAA is synthesized by tryptophan or tryptophan-independent mecha-
nisms [8]. IAA has been found to promote plant growth [9]. It is a tiny molecule with a
Plants 2022, 11, 2559. https://doi.org/10.3390/plants11192559 https://www.mdpi.com/journal/plants