112 Journal of Basic Microbiology 2008, 48, 112 – 117
© 2008 WILEY-VCH Verlag GmbH & Co. KGaA, Weinheim www.jbm-journal.com
Research Paper
Screening of Aspergillus species for occurrence
of lectins and their characterization
Ram Sarup Singh
1
, Ashok Kumar Tiwary
2
and Ranjeeta Bhari
1
1
Carbohydrate and Protein Biotechnology Laboratory, Department of Biotechnology, Punjabi University,
Patiala – 147 002, Punjab, India
2
Department of Pharmaceutical Sciences and Drug Research, Punjabi University, Patiala – 147 002,
Punjab, India
Ten species of Aspergillus were screened for occurrence of lectins. Each of the species was
investigated for the occurrence of extracellular, surface-bound and intracellular lectin ac-
tivities. As many as four species namely, Aspergillus niger, Aspergillus versicolor, Aspergillus rugu-
losus and Aspergillus nidulans, were found to possess intracellular lectin activities, while none of
the species showed extracellular or surface-bound lectin activities. Each of the lectin was
characterized with respect to blood group and carbohydrate specificities. All the lectins were
found to agglutinate human erythrocytes, irrespective of their blood group and pig erythro-
cytes. However, they did not show agglutination with sheep or goat erythrocytes. Of the
various carbohydrates tested, all lectins were found to be specific for inulin, mucin, asia-
lofetuin, N-acetyl galactosamine, melibiose, D-ribose, L-fucose, D-arabinose, D-sucrose and
D-mannitol. The minimum inhibitory concentration of each of the specific sugars was also
determined. The lectins were partially purified using ammonium sulfate precipitation tech-
nique. Each of the lectin was found to be precipitated at 40 – 50% saturation of ammonium
sulfate, yielding about 80% of lectin activity.
Keywords: Lectin / Aspergillus / Haemagglutination / Carbohydrate specificity
Received: October 09, 2007; accepted November 16, 2007
DOI 10.1002/jobm.200700314
Introduction
*
Lectins are polyvalent or univalent oligomeric proteins
of non-immune origin [1]. They bind reversibly with
specific sugars and are capable of precipitating polysac-
charides, glycoproteins and glycolipids bearing specific
sugars and play a role in cell recognition [2]. Lectins
have found many clinical applications such as blood
typing, carriers of chemotherapeutic agents, as mito-
gens and taxonomic markers of specific microorgan-
isms [3]. These have been found to be associated with
murine lymphocyte ‘homing receptor’ MeL 14 [4] and
human leukocyte adhesion molecule ELAM-1 [5]. Lec-
tins bind to specific carbohydrate moieties present on
Correspondence: Dr. R. S. Singh, Department of Biotechnology, Pun-
jabi University, Patiala – 147 002, Punjab, India
E-mail: rssingh11@lycos.com
Phone: +91 175 304 6262
Fax: +91 175 2283073
the apposing cells. This specificity of lectin-carbo-
hydrate binding has been exploited for targeting drugs
or diagnostic agents to normal and diseased tissues. The
role of lectins for drug targeting has been reviewed in
several publications [6, 7].
Lectins are widely distributed in nature and have
been reported in almost all groups of organisms like
viruses, bacteria, fungi, invertebrates, vertebrates and
plants [1]. Amongst the microbes, lectins in bacteria
and protozoa have been extensively studied [8 – 12].
Although there are several reports on fungal lectins
found in fruiting bodies [13 – 19], but only few reports
are there on mycelial lectins [20]. Amongst the fungi
belonging to genus Aspergillus, lectins have been re-
ported in Aspergillus fumigatus [21, 22] and Aspergillus
oryzae [23]. Aspergillus species are highly aerobic and are
found in almost all oxygen-rich environments, where as
a result of the high-oxygen tension, they commonly
grow as molds on the surface of substrates. Aspergillus