Research Article
Enteric Pathogens and Coinfections in
Foals with and without Diarrhea
Giovane Olivo,
1
Thays Mizuki Lucas,
2
Alexandre Secorun Borges,
1
Rodrigo Otávio Silveira Silva,
3
Francisco Carlos Faria Lobato,
3
Amanda Keller Siqueira,
4
Domingos da Silva Leite,
4
Paulo Eduardo Brandão,
5
Fábio Gregori,
5
José Paes de Oliveira-Filho,
1
Shinji Takai,
6
and Márcio Garcia Ribeiro
2
1
Department of Veterinary Clinical Sciences, Universidade Estadual Paulista (UNESP), Botucatu, SP, Brazil
2
Department of Veterinary Hygiene and Public Health, Universidade Estadual Paulista (UNESP), Botucatu, SP, Brazil
3
Department of Preventive Veterinary Medicine, Federal University of Minas Gerais (UFMG), Belo Horizonte, MG, Brazil
4
Department of Genetics, Evolution and Bioagents, Campinas State University (Unicamp), Campinas, SP, Brazil
5
Department of Preventive Veterinary Medicine, University of S˜ ao Paulo, S˜ ao Paulo, SP, Brazil
6
Department of Animal Hygiene, Kitasato University, Towada, Aomori, Japan
Correspondence should be addressed to M´ arcio Garcia Ribeiro; mgribeiro@fmvz.unesp.br
Received 15 September 2016; Accepted 23 November 2016
Academic Editor: Eric N. Villegas
Copyright © 2016 Giovane Olivo et al. Tis is an open access article distributed under the Creative Commons Attribution License,
which permits unrestricted use, distribution, and reproduction in any medium, provided the original work is properly cited.
Diarrhea is a major clinical problem afecting foals up to 3 months of age. Te aim of this study was to identify enteric
microorganisms involved in monoinfections and coinfections and the associated virulence factors in healthy and diarrheic foals.
Diarrheic (D) ( = 56) and nondiarrheic (ND) foals ( = 60) up to three months of age were studied. Fecal samples were analyzed
for identifcation of infectious agents (microbiological culturing, molecular techniques, and microscopic analyses). Escherichia coli
fmH (30% versus 25%), Salmonella spp. (25% versus 7%), Strongyloides westeri (25% versus 25%), Clostridium perfringens type A
(21% versus 10%), E. coli ag43 (20% versus 35%), Strongylus (11% versus 18%), and vapA-positive Rhodococcus equi (5% versus 2%)
were the most frequent enteric pathogens detected in D and ND foals, respectively. Te frequency of toxin A-positive C. perfringens
was signifcantly increased in the D ( = 0.033) compared with the ND animals. R. equi strains harboring virulent plasmids were
also identifed (VapA 85-kb type I and VapA 87-kb type I) in D and ND foals. Coinfections were observed in 46% of the D and 33%
of the ND foals. Our results demonstrate the great diversity of enteric pathogens, virulence factors, and coinfections involved in
enteric infections of foals.
1. Introduction
Diarrhea is one the most common causes of mortality of
neonatal foals, which results in economic losses worldwide
[1, 2]. Until 6 months of life, up to 20% of foals have been
reported to sufer from diarrhea caused by infectious agents
[3]. Te etiology of diarrhea in neonatal foals is complex and
involves infectious agents, management, and facilities, as well
as nutritional, environmental, and physiological conditions
[3–5].
Infectious causes of foal diarrhea predominantly include
bacteria, viruses, and parasitic agents [5–7]. Some studies of
diarrhea in neonatal foals have focused on only one pathogen
[2, 8–11] although it is usually caused by diverse combinations
of enteric infections [1, 5, 7, 12, 13]. Te understanding of
the prevalence and complexity of major pathogens associated
with enteric infections in neonatal foals is limited [14]. Com-
prehensive studies have highlighted the impacts of enteric
coinfections in foals with diarrhea and have identifed major
virulence factors of some pathogens in these infections [1,
5, 7, 13]. However, few studies have evaluated the normal
microbial population of the equine intestine, particularly in
foals [15–17]. Nondiarrheic foals have been suggested to be
potential reservoirs of enteric pathogens, including agents
with zoonotic potential [13, 18]. Te aim of this study was to
identify enteric microorganisms involved in monoinfections
Hindawi Publishing Corporation
BioMed Research International
Volume 2016, Article ID 1512690, 12 pages
http://dx.doi.org/10.1155/2016/1512690