Estuarine, Coastal and Shelf Science 246 (2020) 107017
Available online 28 September 2020
0272-7714/© 2020 Elsevier Ltd. All rights reserved.
Occurrence of enhanced herbivory in the microbial food web of a tropical
estuary during southwest monsoon
P.M. Sooria
a, *
, N. Nandini Menon
a
, R. Ranith
a
, M. Nair
b
, A. Anjusha
b
, A. Shivaprasad
c
,
K. Ajith Joseph
a
, Saramma A. V
d
a
Nansen Environmental Research Centre (India),KUFOS Amenity Centre,Madavana, Kochi, India
b
CSIR-National Institute of Oceanography, Regional Centre, Kochi, India
c
Department of Science and Technology, Govt. of India, New Delhi, 110016, India
d
Cochin University of Science and Technology, Kochi, India
A R T I C L E INFO
Keywords:
Herbivory
Monsoon
Autotrophic picoplankton
Cochin estuary
ABSTRACT
Tropical monsoonal estuaries have a signifcant role in global fshery as they act as rich feeding and breeding
grounds for many commercial species. Food web dynamic studies in these coastal ecosystems are mainly focused
on the classic food chain which assumes that microbial food webs contribute more towards the recycling of
organic carbon and nutrients rather than production. But recently it is observed that phyto-microbial food web
which is largely dependent on pico-autotrophs can contribute signifcantly towards the higher trophic level.
Based on these observations we have analysed phyto-microbial pathway of food web in a highly productive
tropical monsoonal estuary. We assessed the grazer-prey interaction in the microbial food web of Cochin estuary
during spring intermonsoon and southwest monsoon seasons. Our results show that during spring intermonsoon
the food web is dependent more on heterotrophic bacteria, but in southwest monsoon the dependency of grazers
switches over to autotrophic picoplankton. We also analysed the growth rate, grazing rate and carbon contri-
bution of autotrophic picoplankton experimentally during both seasons and found that autotrophic picoplankton
show higher growth rate during southwest monsoon. The higher grazing rate and clearance rate of autotrophic
picoplankton recorded during southwest monsoon indicate the effcient consumption of autotrophic pico-
plankton crop during the season. We could also observe a change in autotrophic picoplankton community
structure between the seasons with the dominance of picocyanobacteria during spring intermonsoon and that of
picoeukaryotes during southwest monsoon. Thus we infer that high freshwater infux into the estuary during
monsoon results in high turbid and low saline (mesohaline to oligohaline) environment which increases the
growth rate of low light adapted autotrophic picoplankton populations and in turn enhances herbivory (phyto-
microbial pathway) in the microbial food web. Apart from protists, autotrophic picoplankton were found to be
widely consumed by many larval forms of mesozooplankton, gastropods, bivalves, rotifers and large sized flter
feeders either as aggregates or as unicellular forms. Thus monsoon enhanced herbivory can have a signifcant
role in the growth and survival of larval forms recruited into the estuary during monsoon season and in turn the
fshery.
1. Introduction
Studies on monsoon associated pelagic food web dynamics have high
socio-economic impact as they give direct insights in to the global
fshery production (Sreeush et al., 2018). Around 70% of fsh landing
that come from the west coast of Indian subcontinent is due to the high
primary production associated with monsoon induced upwelling (Jyo-
thibabu et al., 2015; Sreeush et al., 2018). Most of the hypotheses
proposed to explain the high production during monsoon emphasise the
variability in the linear food chain as it has got a prominent and direct
impact on the higher trophic levels (L´ evy et al., 2007; Roxy et al., 2016;
Praveen et al., 2016; Ezhilarasan et al., 2018; Sreeush et al., 2018).
Nevertheless, a few studies indicate that the role of microbial food web
in the secondary and tertiary production of Indian coast cannot be
ignored (Madhupratap et al., 1996; Nair et al., 1999; Brown et al., 1999).
Both experimental and feld studies reveal that grazers in the
* Corresponding author. Nansen Environmental Research Centre, India.
E-mail address: soorya.mohan1@gmail.com (P.M. Sooria).
Contents lists available at ScienceDirect
Estuarine, Coastal and Shelf Science
journal homepage: http://www.elsevier.com/locate/ecss
https://doi.org/10.1016/j.ecss.2020.107017
Received 10 May 2020; Received in revised form 7 September 2020; Accepted 16 September 2020