International Journal of Ecosystem 2016, 6(2): 35-42
DOI: 10.5923/j.ije.20160602.02
Root Flavonoids of Convolvulus L. Species in
Markazi Province, Iran
Batoul Bahrami
1
, Mitra Noori
2,*
, Amir Mousavi
3
, Ahmad Khalighi
1
, Aliashraf Jafari
4
1
Department of Horticulture, Islamic Azad University, Science and Research Branch, Tehran, Iran
2
Department of Biology, Faculty of Science, Arak University, Arak, Iran
3
Department of Plant Biotechnology, National Institute of Genetic Engineering and Biotechnology, Tehran, Iran
4
Department of Rangeland, Research Institute of Forests and Rangelands, Tehran, Iran
Abstract In order to study the root flavonoids pattern of Convolvulus roots in 11 populations from four species
(C. arvensis, C. commutatus, C. lineatus and C. pilosellaefolius) originated from Markazi Province, Iran, two dimensional
paper chromatography (2-DPC) and thin layer chromatography (TLC) were used. Results indicated that the roots contained
flavonoid sulfates and flavones C and C-/O glycosides, apigenin, chrysin, genistein, kaempferol, luteolin, rhamnetin, rutin,
and vitexin. hesperidin, isorhamnetin, morin, quercetin and tricine was not found in any of the taxa. Vitexin was the most
found flavonoids in root. There was not any aglycone in the studied populations. All of populations had naringenin with the
exception of CBB
5
, CBB
2
and CBB
6
populations. Myricetin was found in all of populations with the exception of CBB
9
(C. commutatus). Populations of CBB
19
(C. arvensis) had the most number of total flavonoids and C. lineatus and CBB
9
(C. commutatus) had the least one.
Keywords Convolvulus, Root, Chemotaxonomy, Chromatography, Flavonoid, Iran
1. Introduction
Flavonoids are found in fruits, vegetables, grains, bark,
roots, stems, leaves and flowers [1, 2]. Root flavonoids play
significant roles in protecting the plants against pests and
diseases, regulating root growth and functions, influencing
different aspects of nitrogen cycle and exerting allelopathic
growth effects. They also constitute an essential source of
Pharmaceuticals [3]. The flavonoid pathway produces a
diverse array of plant compounds with functions in UV
protection, as antioxidants, pigments, auxin transport
regulators, defense compounds against pathogens and during
signaling in symbiosis [4]. Flavonoid synthesis and
accumulation is often very specific for certain cell types. For
example, along the length of a root, flavonoids are often
accumulated at the root tip and in root cap cells. Specific
flavonoid end-products are also localized to specific cell
types where they could have functions in regulating [5, 6].
Within the cell, flavonoids also show specificity for their
location. Flavonoids have been localized to the nucleus, the
vacuole, the cell wall, cell membranes, and the cytoplasm
[7-10]. To date, >10000 flavonoids have been identified in
plants, and their synthesis appears to be ubiquitous in
plants [11]. The flavonoid pathway is one of the best studied
* Corresponding author:
m-noori@araku.ac.ir (Mitra Noori)
Published online at http://journal.sapub.org/ije
Copyright © 2016 Scientific & Academic Publishing. All Rights Reserved
biosynthetic pathways of specialized metabolites.
Flavonoids are phenylpropanoid metabolites, most of which
are synthesized from p-coumaroyl-CoA and malonyl-CoA
and share their precursors with the biosynthetic pathway for
lignin biosynthesis [12]. However, some rare flavonoids are
synthesized from CoA esters of substrates such as cinnamic
acid or dihydro-coumaric acid [13]. Their diversity stems
from the generation of a number of basal flavonoid structures
that include flavones, flavonols, flavan-3-ols, flavanones,
isoflavonoids, isoflavans, and pterocarpans. The flavonoid
skeleton can be modified by glycosylation, malonylation,
methylation, hydroxylation, acylation, prenylation, or
polymerization, leading to the diversity of end-products [14].
These substitutions have important effects on flavonoid
function, solubility, mobility, and degradation [4].
Flavonoids can also be transported within and between cells
and tissues. Within the cell, flavonoids are likely to move
via vesicle-mediated transport or through membrane-bound
transporters of the ABC (ATP binding cassette) or MATE
(multidrug and toxic extrusion compound) families [15].
Flavonoid transport into vacuoles can be achieved by
conjugation of glutathione with flavonoids in the cytoplasm,
followed by ATP-driven transport via glutathione
S-transferasepumps [16, 17, 18]. Long-distance transport of
flavonoids is less well understood but has been
demonstrated in Arabidopsis, where application of
flavonoids to the root or the shoot led to their transport
towards distal tissues [19].
Plant chemosystematics is the application of chemical