©
2007 THE AUTHORS
JOURNAL COMPILATION
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2 0 0 7 B J U I N T E R N A T I O N A L | 1 0 0 , 1 9 9 – 2 0 3 | doi:10.1111/j.1464-410X.2007.06924.x 199
Investigative Urology
PHYTOESTROGENS FROM
BELAMCANDA CHINENSIS AND PROSTATE CANCER CELLS
THELEN
et al.
Phytoestrogens from Belamcanda chinensis regulate the
expression of steroid receptors and related cofactors in
LNCaP prostate cancer cells
Paul Thelen, Thomas Peter, Anika Hünermund, Silke Kaulfuß*, Dana Seidlová-Wuttke†,
Wolfgang Wuttke†, Rolf-Hermann Ringert and Florian Seseke
Department of Urology, *Institute of Human Genetics and †Department Clinical and Experimental Endocrinology,
Georg-August-University, Göttingen, Germany
Accepted for publication 25 January 2007
PT and TP contributed equally to this study
reverse transcription-polymerase chain
reaction to quantify mRNA expression of the
androgen receptor (AR), the AR coactivator
prostate derived Ets transcription factor
(PDEF), NKX3.1, prostate specific antigen
(PSA) and oestrogen receptor- β (ER- β )
compared with the expression of the
housekeeping gene porphobilinogen
deaminase (PBGD). PSA secretion from LNCaP
cells was measured and protein expression of
the AR investigated by Western blot analysis.
RESULTS
Concomitant with a marked decrease of
tumour cell proliferation BCE down-regulated
the expression of the AR, PDEF, NKX3.1 and
PSA. In the same experiments, the expression
of PBGD was unaltered, whereas ER- β
expression increased. Furthermore, AR protein
and PSA secretion were markedly diminished
after treatments with the BCE.
CONCLUSION
BCE, comprising 13 different phytoestrogens,
decreases the expression of the AR and its co-
activator PDEF concomitant with diminished
cell proliferation and PSA secretion. NKX3.1
expression was also reduced by BCE. We
hypothesise that the positive effects of BCE
are initiated by up-regulation of the ER- β , a
putative tumour-suppressor gene.
KEYWORDS
prostate cancer, LNCaP, phytoestrogens,
androgen receptor, oestrogen, proliferation
OBJECTIVE
To investigate the changes in expression
underlying the marked reduction of tumour
growth in vivo , by analysing the effect of
Belamcanda chinensis extract (BCE) on LNCaP
cells in vitro , as phytoestrogens are
chemopreventive in prostate cancer, and in
previous studies we examined the effects of
the isoflavone tectorigenin isolated from B.
chinensis on LNCaP prostate cancer cells, and
a BCE consisting of 13 phytoestrogenic
compounds on tumour-bearing nude mice.
MATERIALS AND METHODS
LNCaP cells were treated with 100, 400 or
1400 μ g/mL BCE; proliferation was assessed
with an Alamar Blue assay. We used real-time
INTRODUCTION
In Western societies prostate cancer is the
most common malignancy in men; in the USA
each year > 230 000, and in Germany > 40 000,
new cases are currently diagnosed. Prostate
cancer is characterized by strong dietary
influences and a long disease latency period.
This long latency affords opportunities for
intervention with therapies that are designed
to delay disease initiation or progression [1].
Diets are regarded as important in the
transformation from latent into more
aggressive prostate cancer, considering that
the frequency of latent prostate cancer is
evenly distributed among populations [2].
Progression from latent stages into clinically
significant prostate cancer is a process that
generally requires several years. During this
period, dietary phytochemicals might have
chemopreventive effects that could slow or
obviate the development of hormone-
dependent cancer. This is supported by
epidemiological evidence that populations
consuming diets rich in soya have lower
incidence of prostate cancer [3].
Several studies have evaluated the
chemopreventive potential of phytochemicals
or phytoestrogens. Phytoestrogens are
polyphenolic nonsteroidal plant compounds
with oestrogen-like biological activity that are
currently under intensive investigation for
their role in human health, and the reasons
for the geographical differences in prostate
cancer incidence rates [4].
Phytoestrogens can act as selective oestrogen
receptor (ER) modulators and they have been
evaluated for potential androgen-blocking
activity [5]. One indicator of androgen-
blocking is the inhibition of androgen-
regulated proteins such as PSA, which is used
as a surrogate marker of disease progression
in clinical studies. PSA secretion and other
actions of androgens are mediated by the
androgen receptor (AR). Chen et al. [6]
showed that the AR is the crucial factor in
the process by which prostate cancer cells
become hormone-refractory and therapy-
resistant. They showed that cancer
progression in hormone-refractory tumours is
controlled by traces of androgen that remain
even under antiandrogen therapy, and that
can be detected by an overexpressed AR.
Moreover, their experiments showed that AR
antagonists under these conditions acted as
AR agonists. This antiandrogen withdrawal
syndrome is shown by a PSA decrease when