Vol.:(0123456789) 1 3
J Insect Conserv (2017) 21:873–883
DOI 10.1007/s10841-017-0026-8
ORIGINAL PAPER
Water availability drives habitat quality for the butterfy Plebejus
argus in a Mediterranean sand dune landscape
Pilar Fernández
1
· David Gutiérrez
2
· Diego Jordano
1
· Juan Fernández Haeger
1
Received: 26 June 2017 / Accepted: 7 October 2017 / Published online: 16 October 2017
© Springer International Publishing AG 2017
(mutualistic ants and nectar sources). As a result, we found
a high spatial congruence in habitat quality between males
and females, and a moderate congruence between adults and
eggs.
Keywords Lasius niger · Lycaenidae · Nectar sources ·
Resource-based habitat · Water table depth
Introduction
The occurrence of species is afected by the area, isola-
tion and quality of habitat patches (Thomas et al. 2001). In
continuous landscapes, in which the spatial arrangement of
habitat (area and isolation) has little efect on the distribu-
tion of a given species, the relative role of habitat quality
is expected to be more important (Mortelliti et al. 2010).
Vegetation or land cover types are frequently used as sur-
rogates of habitat availability, but the appropriate defni-
tion of habitat quality requires more detailed information
on the resources and conditions that are necessary for the
maintenance of the species (Dennis et al. 2003; Mortelliti
et al. 2010). This idea underlies the ‘resource-based habitat
approach’ (Dennis et al. 2003), which has been suggested a
much better perspective for efcient conservation of species
(Dennis et al. 2006).
Key processes involved in survival and reproduction of
an individual should be the most important clues in habitat
use (Sharp et al. 1974; Stefanescu and Traveset 2009). These
clues include diferent abiotic factors like climate, geology,
nutrients, shelter but also biotic interactions (Kubo et al.
2009; Berg et al. 2013; Lawson et al. 2014; Ochoa-Hueso
et al. 2014; Suggitt et al. 2015). In the case of arthropods,
their life cycle comprises several clearly diferentiated stages
that may depend on diferent resources and conditions for
Abstract For terrestrial species, habitat is frequently
defned as physical patches of a certain land cover type in a
matrix of non-habitat. However, the appropriate defnition
of habitat quality requires more detailed information on the
resources and conditions that are necessary for the mainte-
nance of the species (‘resource-based habitat approach’).
In arthropods, their life cycle comprises several clearly dif-
ferentiated stages that may depend on diferent resources
and conditions for successful development; in turn, males
and females may have diferent habitat requirements due
to their diferent behaviours. Here we examine the habitat
quality for males, females and eggs of the butterfy Plebe-
jus argus in a continuous sand dune landscape in southern
Spain. For both sexes, abundance was related to host plant
density (negatively) and distance to heathland vegetation (a
surrogate for higher water availability; negatively for males
and hump-shaped for females); and, for males only, posi-
tively with fower presence. Egg abundance was positively
related to mutualistic ant frequency and fower presence, and
negatively to distance to heathland. Hence, female and male
adults and eggs of P. argus partly difered in their depend-
ence on resources and conditions, although distance to wet-
ter areas of heathland was the common variable for all mod-
els. This suggests that soil humidity is the key variable for
P. argus habitat quality, probably through both direct efects
on individuals and indirect efects on interacting species
* Pilar Fernández
bv2ferop@uco.es
1
Departamento de Botánica, Ecología y Fisiología Vegetal,
Universidad de Córdoba, Campus de Rabanales. Edifcio
C-4, 14071 Córdoba, Spain
2
Área de Biodiversidad y Conservación, Escuela Superior de
Ciencias Experimentales y Tecnología, Universidad Rey Juan
Carlos, 28933 Móstoles, Madrid, Spain