The Laryngoscope
Lippincott Williams & Wilkins
© 2008 The American Laryngological,
Rhinological and Otological Society, Inc.
Functional Assessments of the Rodent
Facial Nerve: A Synkinesis Model
Tessa Hadlock, MD; Jeffrey Kowaleski, BS; David Lo, BS; Roberto Bermejo, PhD; H. Philip Zeigler, PhD;
Susan Mackinnon, MD; James T. Heaton, PhD
Objectives/Hypothesis: Rodent whisker move-
ment has been used as a tool, after facial nerve manipu-
lation, to quantify functional recovery. We have recently
established a method to study functional correlates of
aberrant regeneration of the facial nerve. Our objective
was to establish normative parameters for both sponta-
neous and induced whisking and blinking behavior in a
large group of normal rats.
Study Design: Prospective animal study.
Methods: Eighty animals underwent quantitative
facial movement testing to measure simultaneous vibris-
sal movement and ocular closure for each side indepen-
dently. Right and left C-1 whisker positions were contin-
uously recorded for 5-minute sessions, and changes in
infrared detection corresponding to eye closure were con-
tinuously recorded. Whisking and blinking were elicited
by delivery of olfactory stimuli (10 s scented airflows) and
corneal air puffs. Whisks were counted and analyzed, and
eye closures were counted.
Results: Whisking amplitude, velocity, and acceler-
ation were consistent with literature values. Air puff de-
livery elicited an ipsilateral blink 99% of the time, a
contralateral blink 18% of the time, and changes in or
initiation of bilateral whisking 70% of the time. Olfactory
stimulus delivery prompted a change in whisking behav-
ior 83% of the time, and eye closure 20% of the time.
Conclusions: This study establishes normative
data for assessing cranial nerve VII-controlled facial
movement in four separate facial regions. We demonstrate
the capability and tendency of animals to move their
orbicularis oculi muscles independently of and simulta-
neously with their midfacial muscles. This model pro-
vides an excellent tool for the study of aberrant regen-
eration after facial nerve injury in the rodent.
Key Words: Rodent facial nerve, synkinesis, whisk-
ing, kinematics.
Laryngoscope, 118:1744 –1749, 2008
INTRODUCTION
After facial nerve injury, patients face two dominant
clinical problems. The first is inadequate recovery of func-
tion of the facial musculature, with weak or incomplete
eye closure, and/or a weak smile on the affected side. The
second problem is aberrant regeneration, where axons
reach different muscle targets than their original targets
during the recovery process, and is extremely common
among those facing delayed recovery after facial nerve in-
sult. This phenomenon, termed synkinesis, results in the
involuntary movement of one segment of the face during
attempts at voluntary movement of another. Extensive in-
vestigative effort has been focused on the former issue
(weakness), and progress toward improved overall axonal
traversing of nerve grafts has been steady. The beneficial
role of Schwann cells, graft architecture, and different
growth promoting substances in the regenerating milieu
has been established, and research approaches that com-
bine favorable biological and physical elements in an ideal
neural regeneration conduit seem promising (for review,
see Ref. 1). With emerging technologies that permit time-
lapse observations of motor nerve axonal extension and
branching during regeneration, our understanding of the
factors that underlie regeneration has deepened.
2–6
The latter problem, of axonal misrouting, has been
investigated far less thoroughly, with only a few labora-
tories concentrating on synkinesis after facial nerve in-
jury, and strategies to control it.
7–13
To date, no medical
therapy exists to prevent or diminish the emergence of
synkinesis, despite the fact that patients at high risk for
its development are easy to identify. Because a clinical
window nearly always exists between identifying patients
as “likely to develop synkinesis,” and the actual onset of
From the Department of Otology and Laryngology (T.H., J.K., D.L.),
Massachusetts Eye and Ear Infirmary, Harvard Medical School, Boston,
Massachusetts, U.S.A.; Biopsychology Program (R.B., H.P.Z.), Hunter Col-
lege, City University of New York, New York, New York, U.S.A.; Division
of Plastic and Reconstructive Surgery, Department of Surgery (S.M.), Wash-
ington University, St. Louis, Missouri, U.S.A; and Department of Surgery
(J.T.H.), Massachusetts General Hospital, Harvard Medical School, Boston,
Massachusetts, U.S.A.
Editor’s Note: This Manuscript was accepted for publication May
8, 2008.
Presented at the American Society for Peripheral Nerve Meetings,
Beverly Hills, California, U.S.A., January 9, 2008.
Supported by NIDCR K-08 DE015665-01A2.
Send correspondence to Tessa Hadlock, MD, Department of Otolar-
yngology–Head and Neck Surgery, Massachusetts Eye and Ear Infirmary,
Boston, MA 02114. E-mail: tessa_hadlock@meei.harvard.edu
DOI: 10.1097/MLG.0b013e31817f5255
Laryngoscope 118: October 2008 Hadlock et al.: Normative Rodent Facial Function
1744