Acta Societatis Botanicorum Poloniae
Article ID: 918
DOI: 10.5586/asbp.918
Publication History
Received: 2021-08-17
Accepted: 2022-04-25
Published: 2022-08-15
Handling Editor
Jan Holeksa; Adam Mickiewicz
University in Poznań, Poland;
https://orcid.org/0000-0001-
5138-484X
Authors’ Contributions
FZ designed the study, analyzed
data, and cowrote the paper;
AS edited and reviewed the
manuscript and wrote the initial
draft; GZ carried out data
visualization and searched
scientifc literature; BZ designed
the study, acquired funding,
supervised the research, and
administered the project;
XL assisted in the revision phase
and validation of models;
JT verifed data points and
assisted in the revision; FZ, AS,
and BZ contributed equally to
this research
Funding
This research was funded by the
National Natural Science
Foundation of China (41601091)
and Integrated Scientifc
Investigation of the North-South
Transitional Zone of China
(2017FY100900) (funders had
no role in designing the
experiments).
Competing Interests
No competing interests have
been declared.
Copyright Notice
© The Author(s) 2022. This is an
open access article distributed
under the terms of the Creative
Commons Attribution License,
which permits redistribution,
commercial and
noncommercial, provided that
the article is properly cited.
ORIGINAL RESEARCH PAPER in ECOLOGY
A Multivariate Regression Model to Explain
the Altitudinal Distribution of Timberlines
on the Eurasian Continent
Fang Zhao
1,2
, Abdul Shakoor
1,2
, Gul Zaib
3
, Baiping Zhang
4*
,
Xincan Lan
1
, Jiale Tang
1
1
College of Geography and Environmental Science, Henan University, 475004 Kaifeng,
Henan, China
2
Key Laboratory of Geospatial Technology for the Middle and Lower Yellow River Regions
(Henan University), Ministry of Education, 475004 Kaifeng, Henan, China
3
Institute of Epigenetics and Epigenomics, Yangzhou University, 225009 Yangzhou, China
4
State Key Laboratory of Resource and Environment Information System, Institute of Geographic
Sciences and Natural Resources Research, Chinese Academy of Sciences, 100101 Beijing, China
*
To whom correspondence should be addressed. Email: zhangbp@lreis.ac.cn
Abstract
e altitude of alpine timberline elevation has been considered to correlate with
certain climatic factors. Many related isotherms (e.g., warmest month 10 °C
isotherm) have been proposed to explain the altitudinal distribution of alpine
timberline at the global scale. However, any climatic index actually has a wide
range at the alpine timberline position worldwide. e altitudinal position of the
alpine timberline is related to far more than just one climatic factor. erefore,
we developed a multivariable model for timberline elevation variability by
collecting data from 473 timberline sites on the Eurasian continent. We analyzed
12 climatic variables that potentially account for timberline variation. Principal
component and regression analyses were used to mine four climatic variables.
e mean temperature of the warmest month (MTWM), mean temperature of the
coldest month (MTCM), climatic continentality (K), and annual precipitation
(AP) explained 95% of the variability of timberline elevation. MTWM, MTCM, K,
and AP contributed 18%, 41.28%, 34.9%, and 5.82%, respectively, to the
altitudinal distribution of alpine timberline on the whole continent; 20%, 44%,
28.86%, and 7.14% in the eastern continent; and 17.71%, 39.79%, 40.21%, and
2.29% in the western continent. We showed that MTWM, MTCM, K, and AP are
deterministic factors for the altitudinal distribution of alpine timberline in the
Eurasian continent. MTCM and K contributed to explaining the altitudinal
distribution of timberline both in the entire, eastern, and western parts of the
Eurasian continent. Our research highlights the signifcance of MTCM for the
altitudinal distribution of timberline.
Keywords
Eurasia; timberline elevation; climatic variables; climatic continentality; mean
temperature of the warmest month; annual precipitation; mean temperature of the
coldest month; multivariable model
1. Introduction
e vegetation boundary that divides closed forests and nonforest in the high
mountain area is referred to as the timberline or treeline (Korner & Paulsen, 2004).
Above this line, forest life forms disappear and are replaced by the elfn wood belt,
alpine meadow, or other high mountain scenery (Troll, 1973). Timberline elevation is
related to climatic factors, topography, and even anthropogenic activities (Holtmeier,
2009). Among these factors, climatic factors, including warm-season temperatures,
are deterministic factors and play a vital role in tree growth and survival in the
Acta Societatis Botanicorum Poloniae / 2022 / Volume 91 / Article 918
Publisher: Polish Botanical Society
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