CHEMICAL ENGINEERING TRANSACTIONS
VOL. 38, 2014
A publication of
The Italian Association
of Chemical Engineering
www.aidic.it/cet
Guest Editors: Enrico Bardone, Marco Bravi, Taj Keshavarz
Copyright © 2014, AIDIC Servizi S.r.l.,
I SBN 978-88-95608-29-7; I SSN 2283-9216
MFA of Clostridium Acetobutylicum Pathway: the Role of
Glucose and Xylose on the Acid Formation/Uptake
Francesca Raganati
a
, Alessandra Procentese
a
, Giuseppe Olivieri
a,b
, Maria
Elena Russo
c
, Piero Salatino
a
, Antonio Marzocchella
a
a
Dipartimento di Ingegneria Chimica, dei Materiali e della Produzione Industriali - Università degli Studi di Napoli
Federico II, P.le V. Tecchio 80, 80125 Napoli – Italy
b
Bioprocess Engineering, Wageningen University, P.O. Box 8129, 6700 EV, Wageningen, the Netherlands
c
Istituto di Ricerche sulla Combustione, Consiglio Nazionale delle Ricerche, P.le V. Tecchio 80, 80125 Napoli - Italy
giolivie@unina.it
Acetone-Butanol-Etanol is typically produced during the second stage of batch fermentations of some
Clostridium strains under selected operating conditions: acids are consumed along with the carbon source
and pH increase This contribution reports a MFA regarding Clostridium acetobutylicum DSM 792
fermentation adopting reference sugars (glucose and xylose) as carbon sources. The attention on these
sugars is particularly relevant because they are the main components of hydrolyzed lignocellulosic
biomass. The results have pointed out that the butyrate formation pathway plays a fundamental role both
in the accumulation of butyrate and in butyrate uptake without acetone formation.
1. Introduction
The concerns regarding energy and environmental issues have revalued the interest in the use of biomass
as a renewable energy source. According to this scenario, studies have bloomed in the scientific literature
regarding the production of energy vectors from a wide spectrum of biomass. Solvent-producing clostridia
could produce acetone, butanol, and ethanol (ABE) from several biomasses such as palm oil waste (Lee
et al., 1995), agro-industrial waste(water)s (Raganati et al., 2013), and agricultural crops (Qureshi et al.,
2001). The remarkable features of the butanol – e.g. hydrophobicity, high energy density, storage and
transportation consistent with current structures – make this alcohol a potential substitute and/or
supplement of gasoline (Table 1) (Cascone, 2008; Masiero et al., 2011).
Table 1: Main features of biofuels vs. gasoline
Characteristics Butanol Ethanol Gasoline
Density (kg/m
3
at 20 °C) 810 789 743
Net Calorific value (Mj/kg) 32 27 43
Octane Number 86 100 87
Solubility in water (% at 25 °C ) 9 100 <0.01
ABE is produced during the fermentations of some Clostridium strains - saccharolytic butyric acid-
producing bacteria - under appropriate operating conditions (C. saccharoperbutylacetonicum, C.
acetobutylicum, C. beijerinckii, C. aurantibutyricum). These strains are able to metabolize a great variety of
substrates, pentoses, hexoses, mono-, di- and polysaccharides (Jones and Woods, 1986). Under batch
conditions, the fermentation process of solvent-producing clostridium strains proceeds with the production
of cells, hydrogen, carbon dioxide, acetic acid and butyric acid during the initial growth phase
(acidogenesis) (Jones and Woods 1986). As the acid concentrations increase (pH decrease), the
metabolism of cells shifts to solvent production (solventogenesis) and acidogenic cells – able to reproduce
themselves - shift to the solventogenesis state with a morphological change (Jones and Woods, 1986).
Two different physiological states must be taken into account for Clostridia: one for the acidogenic phase,
and one for the solventogenic phase. During the acidogenesis the acids production ensures high ATP and
DOI: 10.3303/CET1438057
Please cite this article as: Raganati F., Procentese A., Olivieri G., Russo M.E., Salatino P., Marzocchella A., 2014, Mfa of clostridium
acetobutylicum pathway: the role of glucose and xylose on the acid formation/uptake, Chemical Engineering Transactions, 38, 337-342
DOI: 10.3303/CET1438057
337