Vol.:(0123456789) 1 3
International Urology and Nephrology
https://doi.org/10.1007/s11255-017-1748-y
NEPHROLOGY - ORIGINAL PAPER
From bench to the hemodialysis clinic: protein‑bound uremic toxins
modulate NF‑κB/Nrf2 expression
Milena B. Stockler‑Pinto
1,2
· Christophe O. Soulage
3
· Natália A. Borges
1
· Ludmila F. M. F. Cardozo
1
·
Carla J. Dolenga
4
· Lia S. Nakao
4
· Roberto Pecoits‑Filho
5
· Denis Fouque
6
· Denise Mafra
1,2
Received: 31 August 2017 / Accepted: 11 November 2017
© Springer Science+Business Media B.V., part of Springer Nature 2017
Abstract
Purpose Uremic toxins produced by gut microbiota (indoxyl sulfate—IS, p-cresyl sulfate—p-CS, and indole-3-acetic acid—
IAA) accumulate in hemodialysis (HD) patients and exhibit potent infammatory efects. However, the impact of these toxins
on nuclear E2-related factor 2 (Nrf2) and nuclear factor-kappa B (NF-κB) expression in HD patients remains poorly defned.
The aim of this study was to evaluate the association between uremic toxins and Nrf2/NF-κB expression in vitro (RAW 264.7
macrophage-like cells) and in peripheral blood mononuclear cells from HD patients.
Methods Uremic toxins, C-reactive protein (CRP), interleukin-6 (IL-6) and malondialdehyde (MDA) levels were measured
in ffteen HD patients and nine healthy individuals. RAW 264.7 macrophage-like cells were incubated with IS, as a prototype
of protein-bound uremic toxin. Nrf2 and NF-κB expressions were analyzed by RT-qPCR.
Results HD patients presented high levels of infammatory markers, MDA and uremic toxins. In addition, they presented high
NF-κB and low Nrf2 expression. Uremic toxins were positively correlated with NF-κB expression (IS, ρ = 0.58, p < 0.003;
p-CS, ρ = 0.71, p < 0.001; IAA, ρ = 0.62, p < 0.001) and negatively with Nrf2 (IS, ρ = − 0.48, p = 0.01; p-CS, ρ = − 0.46,
p < 0.02). Uremic toxins also exhibited positive correlations with CRP and MDA levels. Multivariate analysis revealed that
p-CS is a determinant factor of NF-κB expression. In RAW 264.7 culture, NF-κB mRNA expression was stimulated by IS,
while Nrf2 was downregulated.
Conclusions Thus, uremic toxins may stimulate NF-κB mRNA and decrease Nrf2 expression in HD patients and, conse-
quently, trigger infammation and oxidative stress.
Keywords Chronic kidney disease · Infammation · Oxidative stress · NF-κB · Nrf2 · Uremic toxins
Introduction
Studies in chronic kidney disease (CKD) patients have
identifed an imbalance of the gut microbiota (the so-called
dysbiosis) as a new factor that may contribute to cardiovas-
cular disease (CVD) [1, 2]. Some uremic toxins produced by
the altered gut such as indoxyl sulfate (IS), p-cresyl sulfate
(p-CS) and indole-3-acetic acid (IAA) are accumulated in
CKD patients and can lead to pathological conditions such
as infammation and oxidative stress which explain in part
the risk of CVD in these patients [3, 4].
IS is an organic anion metabolized in the liver from
indole, produced by the intestinal bacteria as a metabo-
lite of tryptophan derived from dietary proteins [5]. This
toxin is known to exert a wide range of toxic activities
such as endothelial dysfunction and is independently
associated with cardiovascular disease in CKD patients.
Electronic supplementary material The online version of this
article (https://doi.org/10.1007/s11255-017-1748-y) contains
supplementary material, which is available to authorized users.
* Milena B. Stockler-Pinto
milbarcza@gmail.com
1
Graduate Program in Cardiovascular Sciences, Fluminense
Federal University (UFF), Niterói, RJ, Brazil
2
Graduate Program in Nutrition Sciences, Fluminense Federal
University (UFF), Niterói, RJ, Brazil
3
Univ-Lyon, CarMeN Laboratory, Inserm U1060, INSA
Lyon, INRA U1397, Université Claude Bernard Lyon 1,
Villeurbanne, France
4
Departamento de Patologia Básica, Federal University
of Paraná (UFPR), Curitiba, PR, Brazil
5
Pontifícia Universidade Católica do Paraná, Curitiba, PR,
Brazil
6
Department of Nephrology, Centre Hopitalier Lyon Sud,
INSERM 1060, CENS, Université de Lyon, Lyon, France