Flora (2000) 195, 165-171
http://www.urbanfischer.de/journals/flora
©by Urban & Fischer Verlag
Cambial activity and development of wood in Acacia nilotica (L.)
DEL. growing in different forests of Gujarat State
KisHORE S. RAJPUT and K. S. RAo*
* Author for correspondence: Department ofBiosciences, Sardar Patel University, Vallabh Vidyanagar 388120, India
Accepted : September 9, 1999
Summary
Seasonal activity of cambium was studied in Acacia nilotica (L.) DEL. growing in Moist Deciduous (MDF), Dry Deciduous
(DDF) and Scrub land Forests (SF) of Gujarat State for one annual cycle. In MDF cambium was active for major part of the year
reaching peak in August. In DDF, cambial cells underwent divisions throughout the year, but recorded maximum in August and
September. Cell division and differentiation of cambium in SF occurred in two growth flushes. The first flush of cambial activity
began in January, reached peak in April and then declined suddenly in May. The second flush of activity started in October and
continued until November. However, cambium remained inactive during monsoon. Seasonal behaviour of vascular cambium has
been discussed in relation to local climatic conditions and phenology of the trees.
Key words: Seasonal activity, cambium, wood development, Acacia nilotica.
Introduction
Acacia nilotica (L.) DEL. is a moderate sized spiny ever-
green tree indigenous to Sindh, the Deccan, tropical
Africa and is naturalised in all parts of India. The bark
is a major source of tannin and gum-arabic. Wood is also
used in a wide range of artifacts ranging from farming
implements, furniture and good quality timber. A few
studies have been made on seasonal cambial growth
patterns in some species of Acacia (FAHN, 1959, FAHN
et al. 1968). IQBAL & GHOUSE (1987) studied structural
changes in fusiform cambial cells of Acacia nilotica in
relation to age and season. However, no information is
available on seasonal behaviour of the vascular cam-
bium of Acacia nilotica. Therefore, this paper aims to
compare and to understand the response of vascular
cambium of Acacia nilotica, a common evergreen tro-
pical tree growing in Moist Deciduous, Dry Deciduous
and Scrub land Forests.
Materials and Methods
Samples of cambial tissue along with inner bark and
outer sapwood were collected from the main trunk at
breast height of 12-15 year old trees of Acacia nilotica
0367-2530/00/195/02-165 $ 12.00/0
(L.) DEL. having similar trunk diameter (35-40 em)
which were growing in moist deciduous forest at Wag-
hai region of Dangs, dry deciduous forest at Pavagadh
and scrubland forest at Nakhatrana. Periodic collections
were made from January-December 1994 in the second
week of every month. Two trees were sampled each time
to obtain four blocks, no tree being sampled more than
once. The blocks measuring about 60 x 20 mm were
excised with the help of chisel, hammer and single edge
blade and fixed in FAA (BERLYN & MIKSCHE 1976).
Suitably trimmed small pieces of these blocks were sec-
tioned in transverse, radial and tangential longitudinal
planes at a thickness of 12-15 f..Lm on a sliding micro-
tome. After staining with tannic acid-ferric chloride-lac-
moide combination (CHEADLE et al. 1953), the sections
were mounted in DPX.
The term cambium and cambial zone are used to
include the entire population of ray and fusiform cam-
bial cells between the xylem and phloem. Cambial
activity was determined by counting the number of
undifferentiated layers of cambium lying between the
xylem and phloem in transverse section. One hundred
readings of cambial layers at random were selected to
obtain the mean and standard deviation.
Seasonal phenological changes of the trees (such as
sprouting of foliar buds, full foliage, flowering, fruiting
FLORA (2000) 195 165