Snake charming and the exploitation of snakes in Morocco J UAN M. P LEGUEZUELOS ,M ÓNICA F ERICHE ,J OSÉ C. B RITO and S OUMÍA F AHD Abstract Traditional activities that potentially threaten bio- diversity represent a challenge to conservationists as they try to reconcile the cultural dimensions of such activities. Quantifying the impact of traditional activities on biodiver- sity is always helpful for decision making in conservation. In the case of snake charming in Morocco, the practice was in- troduced there  years ago by the religious order the Aissawas, and is now an attraction in the countrys growing tourism industry. As a consequence wild snake populations may be threatened by overexploitation. The focal species for snake charming, the Egyptian cobra Naja haje, is undergo- ing both range and population declines. We estimated the level of exploitation of snakes based on field surveys and questionnaires administered to Aissawas during  , and compared our results with those of a study con- ducted  years previously. Aissawas use four venomous and four non-venomous species for snake charming and we estimate they harvest a minimum of , individuals annually, mostly venomous snakes. For exhibition purposes they selectively remove the largest specimens from the wild (i.e. those that could have the highest reproductive output). Compared to the previous data, we detected () a reduction in the number of species collected, () an increased distance to collecting fields, and () an increase in the market price for snakes, after correction for accumulated inflation, signi- fying a higher demand for these animals. Keywords Aissawas, cobra, Morocco, Naja haje, Sahara, snake charming, tourism, unsustainable use The supplementary material for this article can be found at https://doi.org/./S Introduction I nteractions between people and wildlife have long had an impact on biodiversity. Prehistoric and primitive soci- eties used animals and their derivatives mainly as food but also for clothing, tools, medicine and pets, as well as in magic and religious activities (review in Alves & Rosa, ). Vertebrates, particularly reptiles, have frequently been used for traditional medicine. Alves et al. () iden- tified  reptile species ( families,  genera) currently used in traditional folk medicine, % of which are included on the IUCN Red List (IUCN, ) and/or the CITES Appendices (CITES, ). Among the reptile species being used for medicine, % are snakes. Snakes have always both fascinated and repelled people, and the reported use of snakes in magic and religious activ- ities is global (Alves et al., ). The sacred role of snakes may be related to a traditional association with health and eternity in some cultures (Angeletti et al., ) and many species are under pressure from exploitation as a result (Alves et al., ). The impact of hunting and collecting on snake populations has rarely been studied and remains largely unreported (Gibbons et al., ; Klemens & Thorbjarnarson, ), and declines in snake populations are not receiving the same level of attention as, for example, those in amphibians or marine turtles (Mullin & Seigel, ). Negative effects resulting from exploitation of snake populations must first be assessed before appropriate conservation measures can be established for species of concern. Healers and indigenous peoples have collected wildlife for centuries but in the past harvests were relatively sustain- able and species survival was not threatened (Alvard et al., ). Recent socio-economic-induced changes in human populations, however, such as those related to tourism de- velopment, long considered a clean industry with limited or no negative environmental impacts, are now affecting biodiversity (van der Duim & Caalders, ). One example of this is the population decline in snakes used in snake charming, a practice that began in Morocco c.  years ago when Sidi Mohammed Ben Aissa, an Andalusian sufi, founded the religious order known as the Aissawas in Meknes, central Morocco (Brunel, ). The Aissawas were healers and custodians of knowledge about the local flora and fauna. They used snakes in trad- itional medicine and exhibitions in market places (souks) throughout the country, inspiring the use of snakes in spec- tacles to attract tourists. Snakes used in such spectacles suf- fer from high mortality; when they show obvious signs of ill health they are disposed of and replaced by freshly caught individuals (Highfield & Bayley, ). In  Morocco re- gistered ,, foreign visitors (.% more than during the previous year; Tourisme en Chiffres, ) and tourism JUAN M. PLEGUEZUELOS (Corresponding author) and MÓNICA FERICHE Department of Zoology, University of Granada, E-18071 Granada, Spain E-mail juanple@ugr.es JOSÉ C. BRITO Centro de Investigação em Biodiversidade e Recursos Genéticos, University of Porto, Vairão, Portugal SOUMÍA FAHD Department of Biology, Faculty of Science, Abdelmalek Essaâdi University, Tétouan, Morocco Received May . Revision requested  June . Accepted August . First published online  December . Oryx, 2018, 52(2), 374381 © 2016 Fauna & Flora International doi:10.1017/S0030605316000910 https://www.cambridge.org/core/terms. https://doi.org/10.1017/S0030605316000910 Downloaded from https://www.cambridge.org/core. IP address: 34.228.24.229, on 06 Jul 2020 at 05:53:57, subject to the Cambridge Core terms of use, available at