Research Article
Efficacy of EDTA and Phosphorous on Biomass Yield and
Total Lipid Accumulation in Two Green Microalgae with Special
Emphasis on Neutral Lipid Detection by Flow Cytometry
Gour Gopal Satpati, Prakash Chandra Gorain, and Ruma Pal
Phycology Laboratory, Department of Botany, University of Calcutta, 35 Ballygunge Circular Road, Kolkata, West Bengal 700019, India
Correspondence should be addressed to Ruma Pal; rpalcu@redifmail.com
Received 10 August 2015; Revised 15 December 2015; Accepted 24 December 2015
Academic Editor: Antonio Randazzo
Copyright © 2016 Gour Gopal Satpati et al. Tis is an open access article distributed under the Creative Commons Attribution
License, which permits unrestricted use, distribution, and reproduction in any medium, provided the original work is properly
cited.
Chlorella ellipsoidea and Chlorococcum infusionum, promising microalgae for biodiesel feedstock production, were treated with
ethylenediaminetetraacetate (EDTA) and phosphorous to induce stress which was then followed by fow cytometry to study the
enhanced intracellular neutral lipid content. Treatment resulted in up to a threefold increase in total lipid content of Chlorella
(41.8 ± 1.9% at 16 days of incubation period) and more than twofold increases in Chlorococcum (31.3 ± 1.0% at 18 days of incubation
period) under phosphorous starvation in the culture. It was observed that maximum biomass yields in Chlorella and Chlorococcum
were 1.56 ± 0.06 and 2.17 ± 0.12 g/L at 1.5 g/L of phosphorous afer 20 and 18 days of incubation periods, respectively. Te qualitative
analyses of neutral lipid bodies under stress conditions were performed by confocal microscopy and revealed bright golden-yellow
lipid droplets in stress exposed cells. Signifcant increase of monounsaturated fatty acids under the nutrient limited conditions
was suitable to produce biodiesel. Te maximum biomass (g/L) and lipid content (% dry cell weight) at diferent stresses showed
signifcant results ( < 0.05) by single-factor Analysis of Variance (ANOVA) followed by Duncan’s Multiple Range Test (DMRT).
1. Introduction
Continued exploitation of the world’s fossil fuel reserves is
unsustainable due to their depleted status and simultaneous
steady accumulation of greenhouse gases in the atmosphere.
Development of sustainable energy sources is therefore nec-
essary to overcome this problem. Te production of biodiesel
from photosynthetic microorganisms is considered as an
efective strategy to produce renewable energy. One of the
most promising candidates for production of alternative
energy is microalgae because of their higher growth rate and
oil content than those of the conventional energy crops such
as jatropha, soybean, palm, canola, and corn [1–3]. Addition-
ally, some conditions of cultivation should be strategically
changed to enhance microalgal lipid. Most recent studies have
found that the lipid accumulation in microalgae could be
increased through altering parameters such as temperature,
light, CO
2
, salinity, and nutrient concentration [4–10].
Nutrient is the most important regulator of microalgal
growth and lipid accumulation. Many microalgal species
can synthesize large quantities of lipid and carbohydrates
along with other bioactive molecules under varied nutrient
supplement conditions. Tese newly synthesized lipids and
carbohydrates can be directly converted to biodiesel and
bioethanol, respectively [1, 11, 12]. Under diferent nutrient
starvation, microalgae can accumulate large quantities of
neutral lipid in the form of triacylglycerol (TAG) within
the cell cytoplasm [11]. A few reports were available on the
efect of ethylenediaminetetraacetate (EDTA) on growth and
lipid productivity in microalgae. In Nannochloropsis oculata
the dry weight of the biomass and lipid gradually increased
with increased concentration of EDTA [10]. Te marine
diatom Amphiprora paludosa synthesized 65.64% lipid under
0.026 mM of Fe-EDTA [13].
Simultaneous efect of phosphorous (P) infuences the
growth and lipid synthesis in microalgae [9, 14–17]. Efect
of P defciency in Chlorella vulgaris was studied by Mutlu
et al. [9]. Te lipid accumulation and growth characteristics
of Chlorella kessleri under diferent concentration of P were
Hindawi Publishing Corporation
Advances in Biology
Volume 2016, Article ID 8712470, 12 pages
http://dx.doi.org/10.1155/2016/8712470