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Mol Genet Genomics
DOI 10.1007/s00438-015-1003-0
ORIGINAL PAPER
Genetic diversity and association mapping of bacterial blight
and other horticulturally important traits with microsatellite
markers in pomegranate from India
Nripendra Vikram Singh · Venkata Lakshmi Abburi · D. Ramajayam · Ravinder Kumar · Ram Chandra ·
Kuldeep Kumar Sharma · Jyotsana Sharma · K. Dhinesh Babu · Ram Krishna Pal · Dhananjay M. Mundewadikar ·
Thangasamy Saminathan · Robert Cantrell · Padma Nimmakayala · Umesh K. Reddy
Received: 6 January 2015 / Accepted: 27 January 2015
© Springer-Verlag Berlin Heidelberg 2015
cultivated pomegranate collections from India. Our study
placed all 88 germplasm into four clusters. We identified a
cultivated clade of pomegranates in close proximity to Daru
types of wild-type pomegranates that grow naturally near
the foothills of the Himalayas. Admixture analysis sorted
various lineages of cultivated pomegranates to their respec-
tive ancestral forms. We identified four linked markers for
fruit weight, titratable acidity and bacterial blight sever-
ity. PGCT001 was found associated with both fruit weight
and bacterial blight, and the association with fruit weight
during both seasons analyzed was significant after Bonfer-
roni correction. This research demonstrates effectiveness of
microsatellites to resolve population structure among the
wild and cultivar collection of pomegranates and future use
for association mapping studies.
Keywords Heterozygosity · Population structure ·
Association mapping · Fruit quality · Bacterial blight ·
Pomegranate
Introduction
Pomegranate (Punica granatum L.) of the Lythraceae
family is one of the oldest edible fruit known to humans
and grows naturally in India, Afghanistan and Syria; it
is a native of Iran. The fruit was domesticated around
2000 BC (Rana et al. 2007). Although pomegranate is
widely adapted across the world, the five major produc-
ers of pomegranate are India, Iran, China, the USA and
Turkey (Holland and Bar-Ya’akov 2014; Teixeira da Silva
et al. 2013). India gained importance in cultivation of
pomegranate after 1986; thereafter, it steadily increased
to become first in cultivation at the global level, with
approximately 113 thousand ha under cultivation and
Abstract This genetic diversity study aimed to estimate
the population structure and explore the use of association
mapping strategies to identify linked markers for bacterial
resistance, growth and fruit quality in pomegranate col-
lections from India. In total, 88 accessions including 37
cultivated types were investigated. A total of 112 alleles
were amplified by use of 44 publicly available microsatel-
lites for estimating molecular genetic diversity and popu-
lation structure. Neighbor-joining analysis, model-based
population structure and principal component analysis cor-
roborated the genetic relationships among wild-type and
Communicated by S. Hohmann.
Electronic supplementary material The online version of this
article (doi:10.1007/s00438-015-1003-0) contains supplementary
material, which is available to authorized users.
N. V. Singh · R. Chandra · K. K. Sharma · J. Sharma ·
K. D. Babu · R. K. Pal
ICAR-National Research Center on Pomegranate, Kegaon,
Solapur, Maharashtra 413255, India
V. L. Abburi · T. Saminathan · R. Cantrell · P. Nimmakayala ·
U. K. Reddy
Department of Biology, Gus R. Douglass Institute, West Virginia
State University, Institute, WV 25112-1000, USA
D. Ramajayam
ICAR-Indian Institute of Oil Palm Research, Pedavegi,
West Godavari, Andhra Pradesh 534450, India
R. Kumar
ICAR-Sugarcane Breeding Institute’s Regional Centre,
Karnal, Haryana 132 001, India
D. M. Mundewadikar · U. K. Reddy (*)
MGM’s Institute of Biosciences and Technology, Aurangabad,
Maharashtra 431 003, India
e-mail: ureddy@wvstateu.edu