212 | wileyonlinelibrary.com/journal/wre Weed Research. 2020;60:212–220. © 2020 European Weed Research Society
1 | INTRODUCTION
Efforts to manage invasive weeds are often hampered by insufficient
knowledge of the weed's biology in the exotic range (Jäger et al.,
2009; Bennett et al., 2011). Foliar development and leaf lifespan
are important elements of growth models for individual plants (e.g.
Kikuzawa, 1991). Longer leaf longevity should increase primary pro-
ductivity per tree by increasing the number of mature leaves per tree
that have reached their full photosynthetic capacity (Kikuzawa and
Lechowicz, 2011). Although the capacity to produce large canopies
of actively photosynthesising leaves is typical of successful inva-
sive plants (Rejmanek, 1996), little is known about leaf phenology
of many non-indigenous plant species (Geiger et al., 2011), including
the invasive Schinus terebinthifolia Raddi (Brazilian peppertree).
In this study, we examined leaf phenology of S. terebinthifolia as
part of an effort to develop a biological control programme for this
weed in Florida. Schinus terebinthifolia is a dioecious tree native to
Brazil, Argentina, Uruguay and Paraguay (Ewel, 1986). It was intro-
duced to Florida in the 1840s and successfully colonised 35 Florida
counties after escaping cultivation. It is also invasive in southern
California, southern Arizona, Texas and Hawaii and has naturalised
in over 20 countries (Ewel, 1986; Wunderlin et al., 2016). The Florida
Exotic Pest Plant Council lists S. terebinthifolia as a noxious weed
that poses a threat to native plant species throughout its invaded
Received: 9 July 2019
|
Accepted: 14 February 2020
DOI: 10.1111/wre.12414
ORIGINAL ARTICLE
Foliar lifespan, phenology and seasonal dynamics of the
invasive shrub Schinus terebinthifolia
Brian N. Hogg
1
| Keith Stokes
1,2
| Min B. Rayamajhi
3
| John Geiger
4
| Paul D. Pratt
1
1
Invasive Species and Pollinator Health
Research Unit, USDA-ARS, Albany, CA, USA
2
Cooperative Research, Extension, and
Education Services, Northern Marianas
College, Saipan, Commonwealth of the
Northern Mariana Islands, USA
3
Invasive Plant Research Laboratory, USDA-
ARS, Fort Lauderdale, FL, USA
4
Department of Biological Sciences, Florida
International University, Miami, FL, USA
Correspondence
Brian N. Hogg, USDA-ARS, 800 Buchanan
Street, Albany, CA 94710, USA.
Email: brian.hogg@ars.usda.gov
Funding information
South Florida Water Management District
Subject Editor: Brian Schutte, New Mexico
State University, Las Cruces, USA
Abstract
Schinus terebinthifolia is a dioecious tree native to South America that has become
an invasive weed in Florida, southern California, southern Arizona, Texas and Hawaii
and has been naturalised in over 20 countries. Biological control is considered a viable
long-term control option for S. terebinthifolia because release from natural enemies
appears to be at least partly responsible for its success in Florida. We examined leaf
phenology of S. terebinthifolia over a period of 15 months at five sites in central and
southern Florida to provide information that may help in predicting the impacts of
potential biocontrol agents for this weed. We documented leaf lifespan, the season-
ality of leaf development and abscission and the survivorship of leaves that emerged
during either spring, summer or autumn. Average leaf lifespan was >4.5 months at
all sites, and leaf phenology followed the seasons closely. Although S. terebinthifo-
lia possesses leaves throughout the year, leaf production was greatest from April
to September, and most leaves were abscised in February and March. Spring- and
summer-emerging leaves were also longer-lived than leaves produced during autumn.
These results suggest that leaves of S. terebinthifolia would be most vulnerable to
herbivory during the spring and summer months when newly growing leaf tissue is
most plentiful. Biocontrol agents capable of damaging these tissues during spring/
summer might be an effective means of controlling this invasive weed.
KEYWORDS
Anacardiaceae, leaf phenology, leaf emergence, seasonality, leaf lifespan