Parkinsonism and Related Disorders 20S1 (2014) S68–S72
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Parkinsonism and Related Disorders
journal homepage: www.elsevier.com/locate/parkreldis
Aldehyde dehydrogenase 2 in sporadic Parkinson’s disease
Tanja Maria Michel
a
, Ludwig K¨ asbauer
b
, Wieland Gsell
b
, Julia Jecel
c
, Abigail Jane Sheldrick
a
,
Miriam Cortese
d
, Thomas Nickl-Jockschat
a
, Edna Gr ¨ unblatt
e,f
, Peter Riederer
b,g,
*
a
Department of Psychiatry and Psychotherapy, Rostock University Medical School, Rostock, Germany
b
University Hospital Wuerzburg, Clinic and Policlinic for Psychiatry, Psychosomatics and Psychotherapy, University Wuerzburg, Germany
c
Department of Neurology, Danube Hospital, Center for Social Medicine East, Vienna, Austria
d
RWTH Laboratory for Plastic Surgery, RWTH Aachen University, Aachen, Germany
e
University Clinic of Child and Adolescent Psychiatry, University of Zurich, Zurich, Switzerland
f
Neuroscience Center Zurich, University of Zurich and ETH Zurich, Switzerland
g
Visiting Researcher at Department of Cognitive Brain Sciences, NCGG, Aichi, Japan
article info
Keywords:
Aldehyde dehydrogenase 2
Parkinson’s disease
Mitochondrial defect
Oxidative stress
Aldehyde toxicity
summary
Aldehyde dehydrogenases (ALDH) play a key role in neuronal protection. They exert this function by
metabolizing biogenic amine-related aldehydes, e.g. 3,4-dihydroxyphenylacetaldehyde (DOPAL), and by
protecting neurons against aldehyde- and oxidative stress-related neurotoxicity. The role of these different
isoenzymes has been discussed in other neurodegenerative disorders before. It is somewhat surprising
that only few studies have investigated their role in the aetiology of Parkinson’s disease (PD), in both the
degeneration of dopaminergic neurons and the formation of Lewy bodies. Earlier studies report severe
alterations of the cytosolic isoform of ALDH expression (ALDH 1A1) in the substantia nigra of patients
with PD. However, there are no data regarding the activity of ALDH 2 located at the inner mitochondrial
membrane. Since mitochondrial dysfunctions are hypothesized to be of importance in the aetiology of PD
we have examined the enzymatic activity of mitochondrial ALDH 2 in post-mortem putamen and frontal
cortex of patients with PD and controls. We found that mitochondrial ALDH 2 activity in contrast to the
frontal cortex was significantly increased in the putamen of patients with PD compared to controls.
© 2013 Elsevier Ltd. All rights reserved.
1. Introduction
Idiopathic Parkinson’s disease (PD) is a common neurodegenerative
disorder characterised by a selective degeneration of neuromelanin-
containing dopaminergic (DA) neurons in the pars compacta of
the substantia nigra (SNpc) projecting to the putamen and the
caudate nucleus of the brain. However, the process that leads
to neurodegeneration in PD remains largely unknown. Evidence
suggests that both environmental factors and genetic factors on
the basis of e.g. mitochondrial dysfunctions, oxidative stress (OS)
and aldehyde-related toxicity are major components accounting for
the specific pathophysiology of PD [1]. OS propagates cell death by
increased production of free radicals, which can be either reactive
oxygen species (ROS) or reactive aldehyde species (RAS). A large
number of studies suggest increased OS levels in patients with PD
mediated by mitochondrial dysfunction, decreased concentrations
of antioxidants and elevated iron levels [2]. These processes
contribute to cell degeneration via lipid peroxidation. Oxidative
*Corresponding author. Prof. Dr. Dr. hc. Peter Riederer, University
Hospital Wuerzburg, Clinic and Policlinic for Psychiatry,
Psychosomatics and Psychotherapy, F¨ uchsleinstraße 15, 97080
Wuerzburg, Germany.
E-mail address: peter.riederer@mail.uni-wuerzburg.de (P. Riederer).
deamination leads to the production of aldehydes which may render
toxic if aldehyde dehydroxygenase (ALDH) activity is reduced. In
fact, 4-hydroxy-2-nonenal (4-HNE) and malondialdehyde (MDA)
have been found to be significantly increased in post-mortem SN
of Parkinson’s disease. Both types of free radicals, namely ROS and
RAS, can lead to cellular damage via the modification of proteins and
inhibiting enzymes [3,4]. A significant increase of 4-HNE and MDA
suggests that such aldehydes are not or insufficiently metabolised
into acids by ALDH.
ALDH isoenzymes differ in their affinity to various aldehydes and
show an organ-specific distribution [5]. There are in particular two
ALDH isoenzymes which play a role in detoxifying aldehydes in
the brain, namely ALDH 1A1 and ALDH 2. ALDH 1A1 is located at
the cytosolic site. Therefore it is in close vicinity to monoamine
oxidase (MAO). ALDH 2 is located at the inner mitochondrial
membrane. This demonstrates the importance of getting rid of
toxic aldehydes both within the mitochondria as well as in the
cytosol [6].
There is ample evidence that in PD MAO expression and function
in the SN is not disturbed, while ALDH 1A1 is significantly
reduced or even not present [1,7]. In fact, an increased ratio of
3,4-dihydroxyphenylacetaldehyde (DOPAL) to 3,4-dihydroxyphenyl-
acetic acid (DOPAC) in the putamen of PD suggests decreased
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