Dynamic behavior analysis of carboxymethylcellulose hydrolysis in a chemostat
Rigel V. Gomez-Acata
*
, Pablo A. Lopez-Perez
*
, Rafael Maya-Yescas
**
, Ricardo Aguilar-Lopez
*
*Department of Biotechnology & Bioengineering, CINVESTAV-IPN, No. 2508
D.F. Mexico (Tel: 55-5747-3800; e-mail: raguilar@cinvestav.mx).
** School of Chemical Engineering, Universidad Michoacana de San Nicolas de Hidalgo
Michoacan, Mexico (e-mail: rmayay@umich.mx)
Abstract: At the present time cellulosic waste substrates is a subject of global relevance because is an
alternative source for biofuel production, the complex nature of this substrate and the presence of
microorganisms for its biodegradation may result in complex dynamics which are important to understand
why in many cases could have negative effects in the productivity process. In this paper dynamical
behavior analysis of a chemostat model was carried out for two unstructured kinetic growth models
(Luong and Han-Levenspiel), choosing as study case the carboxymethylcellulose hydrolysis by
Cellulomonas cellulans, where differences related with multiplicity of steady states, hysteresis, periodicity
and unstable operating regions were pointed out.
Keywords: Dynamic behaviors, eigenvalues, hysteresis loop, limit cycle, bifurcation analysis, chemostat
modeling, stability analysis, biological system.
1. INTRODUCTION
Cellulose is the most abundant source of carbon on land; it is
present in the wall of plant cells and is a polysaccharide
consisting of a linear chain of hundreds to thousands β(1→4)
linked D-glucose units. Cellulose is considered the most
important reservoir of carbons to convert glucose, which can
be used for production of ethanol, a promising alternative
energy source for the limited crude oil, organic acids and
other chemicals. (Bo-Kyung et al., 2009; Ye & Jiayang,
2002). Only a small percentage of all microorganisms in the
earth can degrade cellulose principally bacteria and fungi
(Wilson, 2011).
The enzymes produced by these microorganisms have a lot of
applications for example in the textile industry as a detergent
in the pulp and textile industry, for wastewater treatment and
for processing in coffee (Agarwal et al., 2009), reasons for
which it is necessary to produce them in controlled
environments such as bioreactors.
Commonly, bioreactor mathematical models are proposed to
describe the transformations of different substrates by
microorganisms or their enzymes, through setting
phenomenological balances in which unstructured kinetic
growth models are employed to describe the velocity of the
biochemical transformation of substrate to biomass and
metabolites. These unstructured models present disadvantages
like do not take into account the internal structure of the cell
metabolism, biochemical reaction routes nor environmental
effects.
Bifurcation analysis is applied to linear and nonlinear (bio)
mathematical models; it can be used to find optimal operating
conditions and to avoid hazardous conditions for the process
(Namjoshi, Kienle, & Ranmkrishna, 2003). This analysis is
very useful and works revealing all possible equilibrium
points for a given initial operation condition, maintenance free
one or two system parameters in many cases (bifurcation
parameter) such as the dilution rate and initial substrate
concentration. With the above analysis is possible to predict if
the system can present oscillations, hysteresis, steady state
multiplicity and chaos (Alvarez-Ramirez et al. 2009)
(Namjoshi et al. 2003).
The local bifurcation is determined by calculated the system
eigenvalues as in the situation of stability analysis (Gray et al.
1990), it occurs when one of the eigenvalues is close to the
axis of imaginary numbers in the complex plane. The simplest
bifurcations are associated when one of the eigenvalues takes
the value zero (Fold bifurcation) as is the case of the
branching point (BP) and limit point (LP) or when a pair of
conjugate eigenvalues cross the imaginary axis (Hopf
bifurcation, H). The Fold bifurcations are usually the cause of
multiplicity of steady states and hysteresis. Hopf bifurcations
are responsible for the appearance and disappearance of
periodic solutions. (Zhang & Henson, 2001). Furthermore,
this analysis can be used to find optimal operating conditions
and to avoid hazardous conditions for the process.
(Ajbar,2001) (Kuznetzov, 1998), (Namjoshi, Kienle, &
Ranmkrishna, 2003).
In this paper through a dynamic behaviour analysis of the
carboxymethylcellulose hydrolysis by Cellulomonas cellulans
2012 IFAC Conference on Analysis and Control of Chaotic Systems
The International Federation of Automatic Control
June 20-22, 2012. Cancún, México
978-3-902823-02-1/12/$20.00 © 2012 IFAC 132
10.3182/20120620-3-MX-3012.00063