Physicochemical and Chemical Variation in Neem Oils and Some
Bioactivity Leads against Spodoptera litura F.
Jitendra Kumar and Balraj S. Parmar*
Division of Agricultural Chemicals, Indian Agricultural Research Institute, New Delhi 110012, India
Forty-two neem ecotypes of India have been found to show a wide variation in the content of oil,
and their physicochemical characteristics (color, specific gravity, refractive index, iodine value, acid
value, and saponification value), total fatty acid, fatty acid composition (oleic, stearic, palmitic,
linoleic, myristic, arachidic, and behenic acids), and the key meliacins (azadirachtin, nimbin, and
salannin). The azadirachtin content did not correlate with any of the physicochemical and chemical
parameters, but the nimbin and salannin contents correlated significantly with each other. The
refractive index (+ve correlation) and iodine value (-ve correlation) showed weak but significant
correlation with the contents of nimbin and salannin. Insect growth inhibition of Spodoptera litura
revealed a wide variation in the EC
50
of the oils. The salannin and azadirachtin contents of the
oils correlated the most with bioactivity. The iodine and acid values correlated weakly but
significantly with bioactivity.
Keywords: Neem; neem oil; physicochemical variation in neem; chemical variation in neem; neem
bioactivity
INTRODUCTION
The ever increasing emphasis on developing environ-
mentally benign pest control agents has brought neem,
Azadirachta indica A. Juss, to the fore. Several reports
confirming the effectiveness of neem-based products
against various insect pests have appeared in the
literature and can be referred to in the reviews by
Saxena (1989), Schmutterer (1990), Singh (1993), Singh
and Kataria (1991), and others.
Neem oil is a key derivative that finds application as
an important pest control agent. In India, the oil-based
formulations have been prescribed to contain a mini-
mum of 300 ppm of azadirachtin (Parmar and Ketkar,
1993). It is, however, known that besides azadirachtin,
there are several other constituents of the oil that
influence its bioactivity (Schmutterer, 1990).
Neem kernels contain 30-45% (w/w) oil (Koul et al.,
1990; Parmar and Ketkar, 1993). Variation in its
physicochemical characteristics such as color, specific
gravity, refractive index, saponification, iodine and acid
values, and chemical constituents, namely the fatty acid
composition, is reported (Roy and Dutt, 1929; Child and
Ramanathan, 1936; Hilditch and Murti, 1939; Dasa Rao
and Seshadari, 1942; Skellon et al., 1962). The variation
in content of the key meliacin, azadirachtin, is also well
documented (Ermel et al., 1987; Rengasamy et al., 1993).
Despite the variation, specifications for neem oil are
already prescribed (ISI, 1975; The Indian Pharmaco-
poeia, 1966), and the samples for various uses are
required to conform to these.
Even though variation in the various physicochemical
and chemical parameters of the oils has been reported,
it is not known if the content of the meliacins correlates
with any of these parameters. This paper reports
information on this aspect. Additionally, preliminary
information on the effect of these parameters on bio-
activity against Spodoptera litura F. has been gener-
ated.
MATERIALS AND METHODS
Neem Oils. Forty-two neem seed samples from different
areas of India were procured through M/s Neem Mission, Pune,
India. The dry, cleaned seeds were decorticated manually to
obtain kernels which were crushed in a Waring blender and
extracted with n-hexane (60-80 °C) in a Soxhlet extractor for
8 h (at this stage, a drop of hexane extract when evaporated
on a filter paper left no residual oily spot). Hexane was
removed in a rotavapor under reduced pressure at 60 °C to
yield oil.
Chemicals. Standard azadirachtin (purity 95%, HPLC),
M/s Trifolio-M GMBH, Germany, was obtained through the
courtesy of M/s Neem Mission. Reference salannin (purity
85%, HPLC) was obtained from Dr. E. D. Morgan, Keele
University, Staffordshire, U.K., and reference nimbin (purity
85%, HPLC) was obtained from Dr. C. Devakumar of this
Institute. Reference samples of methyl linoleate, methyl
palmitate, and methyl oleate (Aldrich Chemical Co., Milwau-
kee, WI) were obtained through the courtesy of Dr. T. R.
Madan of the Biochemistry Department of this Institute.
Methyl myristate and methyl stearate were prepared in the
laboratory by esterification of their respective acids (Aldrich)
with diazomethane in diethyl ether (AOAC, 1975). For routine
laboratory work, laboratory grade chemicals and solvents were
used. For HPLC analysis, the solvents were of analytical
grade.
Experimental Insect. Twelve-hour-old neonate first in-
star larvae of S. litura F. were used. The culture was
maintained at 27 ( 1 °C, 70-75% relative humidity, and 16:8
h light-dark (LD) and was reared on castor leaves until
transferred to experimental diet (no. 9795, BioServ, Inc.,
French Town, NJ).
Physicochemical Properties. Specific gravity (25 °C),
refractive index (27 °C, M/s Toshniwal India Ltd. refractome-
ter), and iodine, acid, and saponification values (AOAC, 1975)
of oils were determined. The color was examined using
Munsell color charts (1975 edition) by holding 1 mL of the oil
sample in a 3.5 cm × 1 cm diameter glass tube directly behind
the aperture separating the closest chip.
Methyl Esters of Fatty Acids in Neem Oils. Accurate
weights of oils (around 1 g) were saponified with alcoholic
KOH. Alcohol was removed under reduced pressure in a
rotavapor, and the saponified oil was hydrolyzed with dilute
HCl to obtain free fatty acids. The whole content was taken
in a separatory funnel and partitioned with diethyl ether (2
× 5 mL). The ether extracts were esterified with diazo-
methane. The methyl esters were determined by gas-liquid
chromatography (Hewlett-Packard, Model 5890 A, fitted with
coiled glass column, 2 m × 2 mm i.d., packed with 3% SE-30 on
2137 J. Agric. Food Chem. 1996, 44, 2137-2143
S0021-8561(95)00283-4 CCC: $12.00 © 1996 American Chemical Society