Research Journal of Fisheries and Hydrobiology, 3(2): 54-62, 2008
© 2008, INSInet Publication
Corresponding Author: Mohd Hanafi Idris, Department of Animal Science and Fishery, Faculty of Agriculture and Food
Sciences, Universiti Putra Malaysia Bintulu Campus, Nyabau Road, P. O. Box 396, 97008 Bintulu,
Sarawak, Malaysia.
E-Mail: mhanafi@putra.upm.edu.my
54
Biodiversity and Distribution of Pen Shells (Bivalvia: Pinnidae) from the Seagrass
Beds of Sungai Pulai, Peninsular Malaysia
Mohd Hanafi Idris, Aziz Arshad, Japar Sidik Bujang Mazlan Abd. Ghaffar and
1,3 2 1,3 4
Siti Khalijah Daud
3
Department of Animal Science and Fishery, Faculty of Agriculture and Food Sciences,
1
Universiti Putra Malaysia Bintulu Campus, Nyabau Road, P. O. Box 396, 97008 Bintulu,
Sarawak, Malaysia.
Department of Aquaculture, Faculty of Agriculture II, Universiti Putra Malaysia,
2
43400 UPM Serdang, Selangor, Malaysia
Department of Biology, Faculty of Science, Universiti Putra Malaysia, 43400 UPM Serdang,
3
Selangor, Malaysia.
School of Environmental and Natural Resources Science, Faculty of Science and Technology,
4
Universiti Kebangsaan Malaysia, 43600 Bangi, Selangor, Malaysia.
Abstract: Studies on biodiversity and distribution of pen shells were conducted at Merambong
Shoal (N1 19’ 55.62” E103 35’ 57.75”), Tanjung Adang Shoal (N1 19’ 48.03” E103 33’ 59.44”)
o o o o
and Merambong Island (N1 18’ 54.83” E103 36’ 33.37”) off South Western Johor coast, Malaysia
o o
from August 2005 to June 2006. A total of 135 individuals were collected from three different
stations. Pinna bicolor and Pinna atropurpurea were dominance. Higher density of pen shells were
recorded in St 1 with 0.027 ind/m and St 2 was recorded lower density with 0.004 ind/m at
2 2
Merambong Shoal. Univariate analysis at Merambong Shoal population recorded higher values for
diversity and richness indices compared to Tanjung Adang Shoal and Merambong Island but the
value of Evenness index was quite similar between the three stations. Fifty eight individuals of
pen shell were collected from study areas for taxonomy identification. Seven groups of pen shell
were identified base on the internal and external surface of the valves of pen shell. Seven species
were clearly identified which Pinna and Atrina were dominant from the seagrass bed of Sungai
Pulai, Peninsular Malaysia.
Key words: Bivalvia, Pinnidae, Pen Shells, Distribution, Diversity, Malaysia
INTRODUCTION
The Pinnidae or Pen shells are widely distributed
in the Indo-Pacific from southeastern Africa to
Melanesia and New Zealand, north to Japan and to
New South Wales and New Zealand. Pen shells are
also found in Mediterranean and American
waters . Atrina and Pinna species exist as
[19.16.3.25]
metapopulations, composed of small groups or patches
of individuals. Pen shells are generally large bivalves
(30-48 cm long), triangular in shape, thin, shell
tapering to a point and light yellow-brown to dark
brown in color. Pen shells live with their pointed end
embedded in sediment, attached by abundant fine
byssal threads . Pen shells are relatively common
[13,22,8,9]
at the sandy substrate of the seagrass beds (Posidonia
oceanica and Cymodocea nodosa ), lagoons and coral
rubble areas . Most publications of pen shells come
[1,14]
from the researchers in Indo-Pacific, Mediterranean and
American regions . Pen shells show a considerable
[19,16,17]
morphological variation creating taxonomic confusion
for many taxonomists. The genetic heterogeneity
analysis and RAPD marker detection have been used
to identify pen shells up to species level . According
[24]