Citation: Al-Sheboul, S.A.;
Al-Moghrabi, S.Z.; Shboul, Y.;
Atawneh, F.; Sharie, A.H.; Nimri, L.F.
Molecular Characterization of
Carbapenem-Resistant Acinetobacter
baumannii Isolated from Intensive
Care Unit Patients in Jordanian
Hospitals. Antibiotics 2022, 11, 835.
https://doi.org/10.3390/
antibiotics11070835
Academic Editor: Mehran Monchi
Received: 3 June 2022
Accepted: 16 June 2022
Published: 21 June 2022
Publisher’s Note: MDPI stays neutral
with regard to jurisdictional claims in
published maps and institutional affil-
iations.
Copyright: © 2022 by the authors.
Licensee MDPI, Basel, Switzerland.
This article is an open access article
distributed under the terms and
conditions of the Creative Commons
Attribution (CC BY) license (https://
creativecommons.org/licenses/by/
4.0/).
antibiotics
Article
Molecular Characterization of Carbapenem-Resistant
Acinetobacter baumannii Isolated from Intensive Care Unit
Patients in Jordanian Hospitals
Suhaila A. Al-Sheboul
1,
*, Salam Z. Al-Moghrabi
1
, Yasemin Shboul
1
, Farah Atawneh
1
, Ahmed H. Sharie
2
and Laila F. Nimri
1
1
Department of Medical Laboratory Sciences, Faculty of Applied Medical Sciences, Jordan University of
Science and Technology, Irbid 22110, Jordan; szalmoghrabi11@ams.just.edu.jo (S.Z.A.-M.);
yashboul@just.edu.jo(Y.S.); farah.hatawneh@gmail.com (F.A.); nimri01@just.edu.jo (L.F.N.)
2
Faculty of Medicine, Jordan University of Science and Technology, Irbid 22110, Jordan;
ahalsharie16@med.just.edu.jo
* Correspondence: sashboul@just.edu.jo; Tel.: +962-797411589
Abstract: Acinetobacter baumannii is a common cause of healthcare-associated infections (HAI) world-
wide, mostly occurring in intensive care units (ICUs). Extended-spectrum beta lactamases (ESBL)-
positive A. baumannii strains have emerged as highly resistant to most currently used antimicro-
bial agents, including carbapenems. The most common mechanism for carbapenem resistance in
this species is β-lactamase-mediated resistance. Carbapenem-hydrolyzing class D oxacillinases are
widespread among multidrug-resistant (MDR) A. baumannii strains. The present study was conducted
to determine the presence and distribution of bla
OXA
genes among multidrug-resistant A. baumannii
isolated from ICU patients and genes encoding insertion sequence (IS-1) in these isolates. Addition-
ally, the plasmid DNA profiles of these isolates were determined. A total of 120 clinical isolates of
A. baumannii from various ICU clinical specimens of four main Jordanian hospitals were collected.
Bacterial isolate identification was confirmed by biochemical testing and antibiotic sensitivity was
then assessed. PCR amplification and automated sequencing were carried out to detect the presence
of bla
OXA-51
, bla
OXA-23
, bla
OXA-24
, and bla
OXA-58
genes, and ISAba1 insertion sequence. Out of the
120 A. baumannii isolates, 95% of the isolates were resistant to three or more classes of the antibi-
otics tested and were identified as MDR. The most frequent resistance of the isolates was against
piperacillin (96.7%), cephalosporins (97.5%), and β-lactam/β-lactamase inhibitor combinations an-
tibiotics (95.8%). There were 24 (20%) ESBL-producing isolates. A co-existence of bla
OXA-51
gene and
ISAba1 in all the 24 ESBL-producing isolates was determined. In addition, in the 24 ESBL-producing
isolates, 21 (87.5%) carried bla
OXA-51
and bla
OXA-23
genes, 1 (4.2%) carried bla
OXA-51
and bla
OXA-24
,
but all were negative for the bla
OXA-58
gene. Plasmid DNA profile A and profile B were the most
common (29%) in ESBL-positive MDR A. baumannii isolates while plasmid DNA profile A was the
most common in the ESBL-negative isolates. In conclusion, there was an increase in prevalence of
MDR-A. baumannii in ICU wards in Jordanian hospitals, especially those having an ESBL phenotype.
Thus, identification of ESBL genes is necessary for the surveillance of their transmission in hospitals.
Keywords: Acinetobacter baumannii; ICU; multidrug resistance genes; ESBL; plasmid DNA profile;
Oxa carbapenems; insertion sequence; Jordan
1. Introduction
Acinetobacter baumannii is a Gram-negative coccobacillus that is an aerobic, nonmotile,
catalase-positive, and oxidase-negative pathogen. A. baumannii is an opportunistic pathogen
in humans, affecting mainly people with compromised immune systems, and is becoming
increasingly significant as a nosocomial infection [1]. A. baumannii normally inhabits mu-
cous membranes, skin, and soil [1]. The organism is not fastidious in its growth requirement
Antibiotics 2022, 11, 835. https://doi.org/10.3390/antibiotics11070835 https://www.mdpi.com/journal/antibiotics