Accepted Article Norelle L. Daly 1#% , Sunithi Gunasekera 1#^ , Richard J. Clark 1& , Feng Lin 3 , John D. Wade 3 , Marilyn A. Anderson 2 , David J. Craik 1 . 1 Institute for Molecular Bioscience, The University of Queensland, Brisbane, QLD 4072, Australia; 2 Department of Biochemistry, La Trobe University, Melbourne, Victoria 3086, Australia; 3 Howard Florey Institute, University of Melbourne, Victoria 3010, Australia. : David J. Craik, Institute for Molecular Bioscience The University of Queensland Brisbane, QLD 4072, Australia Email: d.craik@imb.uq.edu.au # N.L.D. and S.G. contributed equally to this manuscript. % Current address: Centre for Biodiscovery and Molecular Development of Therapeutics, AITHM, James Cook University, Cairns, QLD Australia. ^ Current address: Department of Medicinal Chemistry, Division of Pharmacognosy, Uppsala University, Box 574, SE 75123 Uppsala, Sweden & Current address: School of Biomedical Sciences, The University of Queensland, Brisbane, Queensland, 4072, Australia. Page 1 of 25 Biopolymers: Peptide Science This is the author manuscript accepted for publication and has undergone full peer review but has not been through the copyediting, typesetting, pagination and proofreading process, which may lead to differences between this version and the Version record. Please cite this article as doi:10.1002/ bip.22977. This article is protected by copyright. All rights reserved.