Taxonomic Note
A new diapsid from the Middle Triassic of southern China
Chun Li,
1
Nicholas C. Fraser,
2
Olivier Rieppel,
3
Li-Jun Zhao,
4
and Li-Ting Wang
5
1
Institute of Vertebrate Palaeontology and Palaeoanthropology, Laboratory of Evolutionary Systematics of Vertebrates, P.O. Box 643, Beijing,
100044, People’s Republic of China 〈lichun@ivpp.ac.cn〉
2
National Museums Scotland, Chambers Street, Edinburgh EH1 1JF, UK 〈nick.fraser@nms.ac.uk〉
3
Field Museum of Natural History, 1400 South Lake Shore Drive, Chicago IL 60605, USA 〈orieppel@fieldmusuem.org〉
4
Zhejiang Museum of Natural History, 71 Jiaogong Road, Hangzhou, Zhejiang 310012, P.R. China
5
Geological Survey of Guizhou Province, Guiyang 550005, P.R. China
Abstract.—The Middle and early Late Triassic of southern China is well known for a remarkable diversity of marine
vertebrates, particularly reptiles, including an abundance of intriguing new forms (e.g., Jiang et al., 2005; Hu et al., 2011;
Li et al., 2016). Here we describe a new diapsid from Yunnan Province. It possesses an elongate neck that exhibits a
remarkable similarity to that of many Protorosauria, yet in other respects the skull and postcranium are much less derived.
The new taxon is part of the so-called Panxian-Luoping Fauna and the deposits correspond to the Upper Member of
the Guanling Formation, comprising thin to medium bedded, gray to dark-gray laminated marly limestone and
limestone, with several layers of bentonite intercalated in the fossil level at Panxian (Wan, 2002; Motani et al., 2008;
Jiang et al., 2009). Their age is Pelsonian (middle Anisian, Middle Triassic) as is indicated by the conodont Nicoraella
kockeli Zone (Sun et al., 2006; Zhang et al., 2009). A recent U-Pb study indicates the absolute age of these middle
Anisian beds to be close to 244 Ma (Wang et al., 2014).
Materials and methods
The description is based on a single, well-preserved, articulated,
and almost complete specimen, including the distal-most caudal
vertebrae (Fig. 1). It was collected from the Guanling Formation
of Luoping County in Yunnan Province. It was prepared by, and
is now housed in, the Institute of Vertebrate Palaeontology and
Palaeoanthropology in Beijing.
Repository and institutional abbreviation.—IVPP, Institute of
Vertebrate Palaeontology and Palaeoanthropology, Beijing, China.
Systematic paleontology
Reptilia
Archosauromorpha von Huene, 1946
?Protorosauria Huxley, 1871
Family incertae sedis
Genus Pectodens new genus
Type species.—Pectodens zhenyuensis n. gen. n. sp. by monotypy.
Diagnosis.—As for type species.
Etymology.—From the Latin pecto meaning to comb and
dens meaning teeth; in reference to the comb-like nature of the
marginal dentition.
Occurrence.—Luoping County of Yunnan Province, China;
Member II of the Guanling Formation, Anisian, Middle
Triassic.
Pectodens zhenyuensis new species
Holotype.—IVPP V18578. Almost complete articulated skeleton.
Diagnosis.—Small, gracile diapsid (about 38 cm long), with long
tail and neck, ten premaxillary teeth, numerous elongate conical
teeth forming comb-like marginal dentition, palatal teeth at least on
the pterygoid present, temporal region short. At least ten, and
probably eleven, or even twelve elongate cervical vertebrae
bearing ribs with a short anteriorly projecting process and a long
posterior shaft that extend parallel to the neck and bridge at least
two intervertebral joints. Dorsals with prominent transverse
processes that terminate in subcircular facets for the ribs. Between
eleven and thirteen dorsal, two sacral, and 41 caudal vertebrae. The
tibia as long, or just slightly longer than, the femur. An elongate
metatarsal V that is not hooked.
Occurrence.—Luoping County of Yunnan Province, China;
Member II of the Guanling Formation, Anisian, Middle
Triassic.
Description.—Positioned on two blocks, the specimen is
preserved with the right side of the skull and neck exposed, but
Journal of Paleontology, 91(6), 2017, p. 1306–1312
Copyright © 2017, The Paleontological Society
0022-3360/17/0088-0906
doi: 10.1017/jpa.2017.12
1306
https://doi.org/10.1017/jpa.2017.12 Published online by Cambridge University Press