Postharvest Biology and Technology, 3 (1993) 235-247 235
© 1993 Elsevier Science Publishers B.V. All rights reserved 0925-5214/93/$06.00
POSTEC 01058
Carbon dioxide action on metabolism of organic
and amino acids in crisphead lettuce *
Dangyang Ke, Mila Mateos * *, Jingtair Siriphanich, Chingying Li
and Adel A. Kader
Department of Pomology, University of California, Davis, CA, USA
(Accepted 7 May 1993)
ABSTRACT
Tissue of crisphead lettuce (Lactuca sativa L.) was kept in air or air enriched with 5% to 20% CO 2
at 0°C for 2 to 9 days followed by transfer to air or the CO2-enriched atmosphere at 20°C for 1 day to
study the mode of CO 2 action on metabolism of organic and amino acids. Exposure to 20% CO 2
reduced the extractable activity of succinate dehydrogenase. The 20% CO 2 treatment caused a
decrease in pH, which partially inhibited the catalytic activity of succinate dehydrogenase and substan-
tially activated glutamate decarboxylase. As a result, succinate and y-aminobutyrate accumulated and
concentrations of malate and glutamate decreased. Such effects on the organic and amino acids were
more pronounced with higher temperature, higher CO 2 concentrations, and longer exposure to CO 2.
The accumulation of succinate and y-aminobutyrate was induced by 15% or 20% CO 2 in all the three
tested cultivars ('Climax', 'Salinas', and 'Winter Haven') and in both green and white tissues of lettuce,
regardless of variation in their susceptibilities to CO 2 injury.
Key words: Lettuce; Lactuca satica; Organic acid; Amino acid; Controlled atmosphere; Physiological
disorder
INTRODUCTION
Crisphead lettuce is very sensitive to CO 2 injury (Stewart and Uota, 1971;
Brecht et al, 1973; Kader and Morris, 1977). Responses of lettuce to elevated CO 2
atmospheres included a decrease in titratable acidity, induction of phenylalanine
ammonia-lyase (PAL), accumulation and oxidation of soluble phenolic compounds,
Correspondence to: Adel A. Kader, Department of Pomology, University of California, Davis, CA
95616-8683, USA.
* Research supported, in part, by a BARD grant no. U.S.-1746-89.
* * The work of M. Mateos was supported by a grant from the Direccion General de Investigation
Cientifica y Tecnica (Ministerio de Educacion y Ciencia), Spain.