Effects of skim milk, skim milk yogurt,
orotic acid, and uric acid on lipid
metabolism in rats
Khursheed P. Navder,* E. Beth Fryer, and Holly C. Fryer
Departments of Foods and Nutrition and Statistics, Kansas State University,
Manhattan, KS, USA
The effects of feeding two milk products (skim milk and skim milk yogurt) and two proposed hypo-
cholesterolemic factors (orotic acid and uric acid) on serum cholesterol (HDL, LDL, total, HDL/
Total and HDL/LDL), liver lipids (total liver lipids and liver cholesterol), and aortal cholesterol were
studied. Ten groups, of nine rats each, were fed isocaloric Chow-based diets containing water, 45%
skim milk (SM), 45% skim milk yogurt (SMY), and 0.0025% orotic acid (OA) or 0.001% uric acid (UA),
without or with cholesterol. The SM diet (with cholesterol) resulted not only in lower total cholesterol
(P < 0.10), LDL cholesterol (P < 0.05), aortal cholesterol (P < 0.01), and liver cholesterol (P < 0.I0),
but also in increased HDL (P < 0.05) and HDL/LDL (P < 0.10) cholesterol ratio. The SMY diet, on
the other hand, resulted in lowered total serum cholesterol (P < 0.05) and aortal cholesterol (P <
0.01) and in higher LDL (P < 0.05) cholesterol. The hypocholesterolemic effects were more marked
for SM than for SMY. Addition of OA and UA to diets increased serum cholesterol, LDL cholesterol,
and total liver lipids; the OA diet also increased liver cholesterol. Neither OA nor UA alone was the
factor responsible for the hypocholesterolemic effects seen with SM and SMY feeding.
Keywords: Skim milk; skim milk yogurt; orotic acid; uric acid; serum lipids; liver lipids; aortal lipid
Introduction
Total serum cholesterol has been identified as a major
risk factor for coronary heart disease. Though many
environmental and genetic variables influence serum
cholesterol, a good correlation exists between total
dietary fat intakes and mean serum cholesterol levels
of different populations. Since dairy products make an
appreciable contribution to saturated fat and choles-
terol content of the diet, whole milk often has been
implicated as a coronary health hazard. However,
Mann and Spoerry, ~ reported a low serum cholesterol
in East African Maasai men, even though their diets
Address reprint requests to Dr. E. Beth Fryer at the Department of
Foods and Nutrition, Justin Hall, Kansas State University, Manhat-
tan, KS 66506 USA.
Supported by Kansas Agricultural Experiment Station Contribution
No. 90-245-J.
*Present address: Department of Nutrition and Food Science,
Hunter College, 425 E 25 Street, New York, NY 10010, USA.
Received April 9, 1990; accepted July 25, 1990.
consisted chiefly of fermented whole milk (as much as
8 1/man/d). That observation has prompted a series of
investigations on the hypocholesterolemic effects of
milk. Various dairy products, such as whole milk, fer-
mented milk, skim milk, yogurt, and buttermilk have
been reported to lower serum cholesterol in man 2"6and
in experimental animals. TM In contrast, little or no hy-
pocholesterolemic effect of milk products was found
in other human ~7-2°and animal studies. 2tz3
Even those researchers who have confirmed the
presence of a cholesterol-lowering effect are not cer-
tain about the responsible factors. Several milk con-
stituents such as orotic acid, 24-29 uric acid, 3°,31 cal-
cium, 3'5'12'32 lactose, 33 milk fat globule membrane, 6 and
products of bacterial fermentation 1°,13 have been sug-
gested.
This study was undertaken to determine the effects
of two dairy products (skim milk and skim milk yo-
gurt) and two proposed hypocholesterolemic factors
(orotic acid and uric acid) on the cholesterol metabo-
lism in normal and experimentally induced hypercho-
lesterolemic rats. In an effort to ascertain the mecha-
nism(s) responsible for the dairy product effect, serum
640 J. Nutr. Biochem., 1990, vol. 1, December © 199o Butterworth-Heinemann