Vol.:(0123456789) 1 3
Conservation Genetics
https://doi.org/10.1007/s10592-020-01307-0
RESEARCH ARTICLE
Detecting genomic variation underlying phenotypic characteristics
of reintroduced Coho salmon (Oncorhynchus kisutch)
Rebekah L. Horn
1
· Cory Kamphaus
2
· Keely Murdoch
2
· Shawn R. Narum
1
Received: 9 December 2019 / Accepted: 19 August 2020
© Springer Nature B.V. 2020
Abstract
For species that have been extirpated from parts of their range, conservation managers often reintroduce individuals to these
areas in hopes of restoring populations to pre-decline conditions. Coho salmon (Oncorhynchus kisutch) have been extirpated
since the early 1900s in the interior reaches of the Columbia River watershed. Starting in the late 1990s, the Columbia River
Treaty tribes were successful in starting a re-introduction program that has established an upper Columbia River Coho
salmon stock. Fish are preferentially selected for broodstock to incorporate naturally occurring phenotypic characteristics
to facilitate local adaptation. On the Wenatchee River in Washington, broodstock are preferentially selected at a lower and
upper river dam, however, only ~ 32% of fsh successfully ascend a 15 km high-gradient reach to the upper river dam. Fish
that successfully ascend the reach generally arrive early in the season and have a better overall body condition. In other
salmonids, phenotypic traits such as return timing has been shown to be under genetic control. To determine if there are
genomic regions that underly the phenotypic traits found to impact migration success up a high-gradient reach, low-coverage
whole genome re-sequencing (lcWGR) was performed on adult fsh returning to the system. Genome-wide association tests
revealed three genomic regions that are associated with fsh return location. Results of the lcWGR suggest that candidate
markers can be incorporated as a genetic screening tool during broodstock selection to preferentially breed fsh that have the
phenotypic characteristics that confer greater potential for steeper and longer migration distances.
Keywords Coho salmon · Pool-seq · GWAS · Reintroduction · Fisheries management
Introduction
Conservation management of species often requires the
movement and reintroduction of individuals into regions
previously extirpated (Seddon et al. 2007). Success of these
programs can be hard to estimate, with the ultimate goal for
populations to reach their carrying capacity and maintain
viable population sizes (Robert et al. 2015). Most reintro-
duction projects focus on mammals or birds with only 4% of
projects focused on fsh (Seddon et al. 2007). In the Pacifc
Northwest, almost a third of the salmonid populations have
been lost since the arrival of Europeans, with most of those
in the interior reaches (Gustafson et al. 2007). Dams operat-
ing along the Columbia and Snake Rivers contributed to the
decline of upriver stocks of Chinook salmon (Oncorhynchus
tshawytscha) (Schaller et al. 1999) and led to the extirpation
of interior river Coho salmon (O. kisutch) (Nehlsen et al.
1991). In salmonids, re-introduction programs are becoming
more numerous to return fsh to habitat no longer volitionally
accessible due to dams or other man-made barriers (Bosch
et al. 2007; Anderson et al. 2014; Galbreath et al. 2014; Sard
et al. 2015; Kozfkay et al. 2019; Matala et al. 2019).
There is considerable diversity in salmonids with respect
to their life history strategies which includes diferential
migration timing, maturation rate, return timing (Smith
et al. 2014), and return distance and/or location (Bennett
et al. 2015; Keefer et al. 2018). This diversity is critical
to understand when implementing reintroduction programs
for salmonids to increase survival of transplanted fsh and
preserve the underlying life history variations (Anderson
Electronic supplementary material The online version of this
article (https://doi.org/10.1007/s10592-020-01307-0) contains
supplementary material, which is available to authorized users.
* Rebekah L. Horn
rhorn@critfc.org
1
Columbia River Inter-Tribal Fish Commission, Hagerman,
ID, USA
2
Yakama Nation Fisheries, Toppenish, WA, USA