[CANCER RESEARCH 50, 3743-3747, June 15, 1990]
Melanocytotoxicity and Antimelanoma Effects of Phenolic Amine Compounds in
Mice in Vivo1
Frank Alena, Kowichi Jimbow,2 and Shosuke Ito3
Division of Dermatology and Cutaneous Sciences, University of Alberta, Edmonton, Canada T6G 2C2
ABSTRACT
A phenolic amine compound, 4-S-cysteaminylphenol (4-S-CAP), is a
potent depigmenting agent. To develop more efficacious antimelanoma
agents, we synthesized four homologues of 4-S-CAP: W-acetyW-S-CAP
(W-Ac-4-S-CAP), a-methyl-4-S-CAP, 4-S-homo-CAP, and N,N'-di-
methyl-4-S-CAP. We tested these five compounds in mice in vivo. After
s.c. or i.p. injection of saline solution (in control groups) or one of the
compounds, follicular melanocytes were examined by light and electron
microscopy to assess the degree of melanocytotoxicity; /V-Ac-4-S-CAP
induced the most depigmentation (98%), whether given ¡.p. or s.c. After
injection of 4-S-CAP or W-Ac-4-S-CAP, the number of murine B16F10
melanoma colonies formed in the lungs was determined; 4-S-C\P and
A-Ac-4-S-CAP were almost equally effective, reducing the colonies to 32
and 25% of mean control, respectively. Metabolic studies of the urine
showed 9% of 4-S-CAP and 20% of 7V-Ac-4-S-CAP injected i.p. were
excreted unchanged in 24 h; 1.3% of the A'-Ac-4-S'-CAP was excreted as
4-S-CAP, indicating some conversion. We conclude that /V-Ac-4-S-CAP
is a suitable model for developing chemotherapy to treat melanoma
characterized by high tyrosinase activity and melanin synthesis.
INTRODUCTION
Melanin pigment is synthesized solely within melanocytes
and melanoma cells. The enzyme tyrosinase (EC 1.14.18.1)
converts the amino acid tyrosine to L-DOPA and thence to
dopaquinone, which gives rise to a melanin polymer. This
process occurs in a specific secretory granule, the melanosome
(1, 2). There have been several attempts to use this unique
melanin pathway as a basis for developing chemotherapeutic
agents selectively toxic to melanoma cells, in most cases based
on L-DOPA and related catechols (3-7), but chemical instability
and severe toxicity due to nonspecific oxidation have precluded
success. Because phenolic compounds seem to possess fewer
drawbacks and appear more promising for use in chemotherapy
to combat melanoma, we developed a new subgroup of phenols,
combining phenols with cysteine and cysteamine to yield cys-
teinylphenol and cysteaminylphenol, respectively (8).
Both 4-5-CP4 and 4-5-CAP are substrates for mammalian
tyrosinase, can depigment black hair, and possess an anti-
melanoma property in vivo, whereas their 2-5-isomers are not
tyrosinase substrates and evidence no melanocytotoxic effects
(8). Most importantly, 4-5-CAP has a significant antimelanoma
effect (9). Padgette et al. (10), however, in developing hypoten-
sive agents discovered that 4-5-CAP is a substrate of plasma
MAO. Our earlier study confirmed this result in vitro and
Received 7/25/89; revised 1/30/90.
The costs of publication of this article were defrayed in part by the payment
of page charges. This article must therefore be hereby marked advertisement in
accordance with 18 U.S.C. Section 1734 solely to indicate this fact.
1This study was supported by a grant from the National Cancer Institute of
Canada.
2To whom requests for reprints should be addressed.
3 Professor of Chemistry at the School of Hygiene, Fujita-Gakuen Health
University, Toyoake, Aichi, Japan; Visiting Professor of the Alberta Heritage
Foundation for Medical Research during this study.
'The abbreviations used are: 4-S-CAP, 4-5-cysteaminylphenol; Ar-Ac-4-5-
CAP, A'-acetyl-4-5-cysteaminylphenol; a-Me-4-S-CAP, a-methyl-4-5'-cysteami-
nylphenol; 4-S-HomoCAP, 4-5-homocysteaminylphenol; N,N-D\Me-4-S-CAP,
A',A''-dimethyl-4-5-cysteaminylphenol; HPLC, high performance liquid chroma-
tography; L-DOPA, L-dihydroxyphenylalanine; MAO, monoamine oxidase;
PTCA, pyrrole-2,3,5-tricarboxylic acid.
indicated that 4-5-CAP could produce a cytotoxic aldehyde
through interaction with plasma MAO (11). To overcome these
difficulties, we synthesized four homologues of 4-5-CAP; N-
Ac-4-S-CAP, a-Me-4-S-CAP, 4-5-homo-CAP, and N,N-DiMe-
4-5-CAP. We found that all four compounds are substrates for
tyrosinase and that 4-5-HomoCAP is also a substrate for MAO
(12).
The depigmentation potency in black follicles of test com
pounds given s.c. is a reliable screening method for evaluating
melanocytotoxicity (13). In the present study, in mice in vivo
we compared this property (visible change and chemical mela
nin content) of the four new phenolic amine compounds and
the mother compound, 4-5-CAP, given both i.p. and s.c., and
assessed the antimelanoma effect of 4-5-CAP and the most
efficacious depigmenting compound by assaying the formation
of melanoma colonies in the lungs.
MATERIALS AND METHODS
Animals. Breeding pairs of C57BL/6J black mice were purchased
from The Jackson Laboratory (Bar Harbor, ME), and their female
progeny were used in the study when 8 weeks old and weighing 17.0 g.
All procedures were approved by the University's Health Sciences
Animal Welfare Committee.
Chemicals. The methods used to synthesize the phenolic amine
compounds [4-5-CAP, W-Ac-4-S-CAP, a-Me-4-S-CAP, 4-S-Homo-
CAP, and A',A'-DiMe-4-5-CAP] were as reported previously (8, 9, 11).
Briefly, the basic compound, 4-S-CAP, was prepared through refluxing
a mixture of phenol and cysteamine with IIBr (Fig. 1). In all experi
ments, the agent injected into controls was l N saline solution.
I .DSU. The LD50 for each compound (n = 6 mice each) was established
by a single i.p. injection of the test drug dissolved in normal saline
solution, in a dose range of 100 to 1300 mg/kg body weight. The LD50
was 600 mg/kg for 4-5-CAP, 1200 mg/kg for W-Ac-4-S-CAP, 500 mg/
kg for a-Me-4-S-CAP, 350 mg/kg for 4-S-HomoCAP, and 300 mg/kg
for N,N-DiMe-4-S-CAP.
Melanoma Cell Line. The murine B16F10 melanoma cell line, with
a strong metastatic property to form tumor colonies in lungs, was
kindly supplied by Dr. B. M. Longenecker (Department of Immunol
ogy, University of Alberta). Cells were grown in T-75 flasks in Dulbec-
co's modified Eagle's medium (GIBCO, Grand Island, NY) supple
mented with 10% fetal calf serum (GIBCO), penicillin (100 units/ml),
and streptomycin (100 jig/ml). Cells were incubated at 37°Cin a
humidified atmosphere of 5% CO2 in air.
Melanocytotoxicity in Vivo. Black hairs were plucked manually from
the back of the mice, to initiate new anagen growth and activate
follicular melanocytes with increased tyrosinase activity. Thirty mice
were randomized into six groups of five (one control and one for each
compound). Starting on day 1, daily for 14 days the agent was injected
i.p. or was infiltrated s.c. in an area where hair follicles had been
plucked; the dose was 300 mg/kg body weight.
Melanin Content. In the same 30 mice, hair follicles were harvested
on day 22 (early telogen phase) and their eumelanin content was
analyzed. Details of the assay, including chemical degradation of mel
anin and HPLC, were as before (14). Briefly, a 10-mg hair sample is
homogenized in water at a concentration of 10 mg/ml; the 200-^1
homogenate is transferred to a screw-capped test tube, mixed with l M
HjSO4 (800 n\), and oxidized with 3% KMnO4. The product, PTCA,
is analyzed by HPLC with a UVL detector; samples are measured in
duplicate. The eumelanin content is expressed as PTCA (ng/mg) versus
3743
Research.
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