Andrologia. 2019;00:e13313. wileyonlinelibrary.com/journal/and
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https://doi.org/10.1111/and.13313
© 2019 Blackwell Verlag GmbH
1 | INTRODUCTION
Type 2 diabetes mellitus is a kind of metabolic disease that results
from defects in insulin secretion, insulin action or both and is charac‐
terised by hyperglycaemia (Association, 2014). Indeed, hyperglycae‐
mia is considered to generate reactive oxygen species through both
enzymatic and nonenzymatic pathways that are associated with
damage and dysfunction of different organs such as male repro‐
ductive organ (Fakhruddin, Alanazi, & Jackson, 2017). It is believed
that spermatozoa are vulnerable to reactive oxygen species damage
(Agarwal, Virk, Ong, & Plessis, 2014; Amaral et al., 2013).
Metabolism of glucose is an important event in spermatogen‐
esis. Many studies confirmed the detrimental effect of diabetes
on semen parameters, chromatin integrity and DNA fragmenta‐
tion in human and other mammals. Most of these changes are
oxidative in nature (Bahmanzadeh, Vahidinia, Mehdinejadiani,
Shokri, & Alizadeh, 2016; La Vignera, Condorelli, Vicari, D'Agata,
& Calogero, 2012; Maiorino, Bellastella, & Esposito, 2014). It is
Received: 30 December 2018
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Revised: 7 April 2019
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Accepted: 17 April 2019
DOI: 10.1111/and.13313
ORIGINAL ARTICLE
Resveratrol supplementation improves DNA integrity and
sperm parameters in streptozotocin–nicotinamide‐induced
type 2 diabetic rats
Maryam Bahmanzadeh
1,2
| Mohammad Taghi Goodarzi
3
| Azam Rezaei Farimani
4
|
Nasibeh Fathi
2
| Zohreh Alizadeh
1,2
1
Endometrium and Endometriosis Research
Center, Hamadan University of Medical
Sciences, Hamadan, Iran
2
Department of Anatomical sciences, School
of medicine, Hamadan University of Medical
Sciences, Hamadan, Iran
3
Department of Biochemistry, Shahrood
Branch, Islamic Azad University, Shahrood,
Iran
4
Department of Basic Medical
Sciences, Neyshabur University of Medical
Sciences, Neyshabur, Iran
Correspondence
Zohreh Alizadeh, Department of Anatomical
sciences, Hamadan University of Medical
Sciences, Shahid Fahmideh Ave P.O.Box.
65178‐518, Hamadan, Iran.
Emails: alizadeh@umsha.ac.ir; alizadeh.
zohreh@gmail.com
Funding information
Vice‐chancellor for Research and
Technology, Hamadan University of
Medical Sciences, Grant/Award Number:
9311145896
Abstract
Reproductive dysfunction is one of the diabetes complications. Resveratrol, a poly‐
phenol compound, shows antidiabetic and antioxidant effects. The aim of the present
study was to investigate the protective effects of resveratrol on sperm parameters
and chromatin quality in experimentally induced type 2 diabetes by streptozotocin
and nicotinamide. Forty male adult Wistar rats were grouped into normal control,
diabetic control and resveratrol‐treated diabetic groups (1, 5 and 10 mg/kg orally
treated for 30 days). Type 2 diabetes was induced using a single dose of streptozo‐
tocin and nicotinamide by intraperitoneal injection. Then, the different parameters
and chromatin condensation of the epididymal extracted spermatozoon were studied
using aniline blue (AB), acridine orange (AO) and toluidine blue (TB) staining. The
sperm parameters including count, motility and viability had significant reduction in
diabetic rats ( p < 0.05). Resveratrol increased count, motility and viable spermatozoa
relative to the diabetic group ( p < 0.05). The mean percentage of AB, AO and TB
staining positive spermatozoa was increased in diabetic groups compared to control
( p < 0.001) and decreased after treatment with 1 and 5 mg/kg resveratrol ( p < 0.001).
The results of AO and TB staining showed that resveratrol did not have any beneficial
effect on chromatin condensation and denatured DNA at the dose of 10 mg/kg.
KEYWORDS
antioxidants, chromatin, diabetes mellitus, hyperglycaemia, male reproductive system,
resveratrol, spermatozoon