Prostaglandins, Leukotrienes and Essential FattyAcids (1998) 59(1 ), 55-62
© HarcourtBrace& Co. Ltd 1998
Systematic pharmacological
approach to the characterization of
NSAIDs
M. Lora, ~ J.-B. Denault, ~ R. Leduc, ~ A. J. de Brum-Fernandes ~'2
~Department of Pharmacology Faculty of Medicine, Universite de Sherbrooke, Canada
2Department of Medicine (Rheumatic Diseases Unit), Faculty of Medicine, Universit6 de Sherbrooke, Canada
Summary Non-steroidal anti-inflammatory drugs (NSAIDs) are widely used for the treatment of inflammatory
diseases. NSAIDs inhibit cyclooxygenase (COX), the rate limiting enzyme responsible for the conversion of
arachidonic acid into prostaglandins. Recent studies have shown the existence of two isoforms of cyclooxygenase:
COX-l, now often referred to as the constitutive form, and COX-2, an inducible form which is the major isoenzyme
involved in prostaglandin synthesis in inflammation and other pathological situations. Since inhibition of prostaglandin
production in tissues where they play a physiological role leads to important side effects, a COX-2 preferential inhibitor
would present therapeutical advantages. In the present study, we evaluated the inhibitory properties of
cyclooxygenase inhibitors on human COX-1 and COX-2 using a heterologous expression system. We investigated
instantaneous inhibition and pre-incubation inhibition as well as time recovery of cyclooxygenase activity assays with
the aid of four NSAIDs: mefenamic acid, indomethacin, aspirin and NS-398. Our results demonstrate that
instantaneous inhibition assays have little correlation with clinical results. Inhibition assays using pre-incubation with
the drugs tested, however, more closely resemble the data from in vivo studies. Cyclooxygenase recovery assays
enabled better characterization of simple competitive inhibitors, competitive reversible time-dependent inhibitors and
irreversible time-dependent inhibitors. The data illustrate the usefulness of our system in allowing a better
determination of the pharmacological characteristics of NSAIDs as well as permitting a comparison among different
drugs.
INTRODUCTION
Cyclooxygenase (COX §, E.C. 1.14.99.1) is the first enzyme
involved in the synthesis of prostaglandins, catalyzing the
formation of prostaglandin H2 (PGH2) from arachidonic
acid. The unstable PGH2 is the common precursor of
prostaglandins and thromboxane. Cyclooxygenase (COX)
is the primary target of non-steroidal anti-inflammatory
drugs (NSMDs), and this action is believed to be respon-
sible for their anti-inflammatory action. 1 Unfortunately,
inhibition of prostaglandin production in organs such as
the stomach and kidney, where they play important
physiological roles, 2-3 can result in gastric lesions and
nephrotoxicity, important side effects that limit the use
of these drugs.
Received 15 October 1997
Accepted 17 December 1997
Correspondence to: Artur J. de Brum-Fernandes, MD, PhD, 3001, 12th
Avenue Nord, Rheumatic Diseases Unit, Sherbrooke, PQ, Canada, J1H 5N4.
Tel: 819 564 5261; Fax: 819 569 5265; E-mail: a.fernan@courrier.usherb.ca
Recent studies have shown the existence of two isoforms
of COX. The first isoenzyme described, cyclooxygenase-1
(COX-I) is constitutive and produces prostaglandins
which have physiological functions such as protecting
the stomach and kidney from damage. 2-3 A more recently
discovered isoenzyme, cyclooxygenase-2 (COX-2) is
induced by inflammatory stimuli such as cytokines 4-8 a nd
produces prostaglandins which contribute to the pain
and swelling of inflammation.
The cDNAs encoding COX-1 and COX-2 have been
cloned from mouse 9-11 as well as human sources 12-15and
expressed in vitro. 1°<3,~6<9 The differences in primary
structures between COX-1 and COX-2 suggest that this
could lead to divergent affinities for fatty acid substrates
and NSAIDs. It has been shown that murine COX-1 and
COX-2 differ in their sensitivity to inhibition by some
NSAIDs, 1~ as do human COX-1 and COX-2. 2° In this
context, selective inhibition of COX-2 could provide an
anti-inflammatory action without the side-effects of the
currently available NSMDs.11,2>22
55