Combining the biological nitrogen and sulfur cycles in
anaerobic conditions
F. Fdz-Polanco, M. Fdz-Polanco, N. Fernández*, M.A. Urueña, P.A. García and S. Villaverde
Department of Chemical Engineering, Valladolid University, 47011 Valladolid, Spain
* Instituto Superior Politécnico José Antonio Echevarría, La Habana, Cuba
Abstract The biochemical processes involved in the anaerobic degradation of carbon, nitrogen and sulfur
compounds can be represented by an oxidation-reduction or electron donor-acceptor scheme. The theoretic
values of Gibbs free energy (DG
0
) calculated from thermodynamic data indicate the feasibility of the
reactions. The interactions C-S and C-N are well known but there is a lack of information about the
interaction N-S. The anaerobic transformation of nitrates using reduced sulfur compounds can be explained
considering that nitrate acts as electron acceptor while reduced sulfur compounds are the electron donors.
A new N-S interaction in anaerobic conditions (ORP = –425 mV) has been experimentally observed when
treating industrial wastewater rich in organic nitrogen and sulfate. The mass balances of the different
nitrogenous and sulfur compounds in the liquid and gas phases clearly indicated an uncommon evolution. An
important percentage of the nitrogen entering the reactor as TKN was removed from the liquid phase
appearing as N
2
in the gas phase. Simultaneously, only part of the sulfate initially present in the influent
appeared as sulfide in the effluent or as hydrogen sulfide in the gas. These experimental observations may
suggest a new anaerobic N-S biological interaction involving simultaneous anaerobic ammonium oxidation
and sulfate reduction, ammonium being the electron donor and sulfate the electron acceptor.
Keywords Anaerobic nitrogen removal; anaerobic sulfur removal; nitrogen/sulfur interaction
Introduction
The biological cycles of carbonaceous organic matter, sulfurous compounds and nitroge-
nous compounds are well established. Anaerobic treatment of industrial wastewater under
conventional conditions generates as end products of the anaerobic degradation: methane
and carbon dioxide for the organic matter, sulfide for the oxidised sulfur compounds and
ammonia for the organic nitrogen compounds (Lettinga, 1995).
All the biochemical processes involved can be represented by an oxidation-reduction or
electron donor-acceptor scheme. The value of the standard Gibbs free energy (DG
0
) of a
chemical reaction can be calculated following the electron transfer scheme and using the
values of DG
0
of the half-reactions. The DG
0
of the half-reactions can be calculated by sim-
ple arithmetic as the difference between standard Gibbs free energies of formation (DG
0
f
)
of products and reactants.
Thermodynamic data are compiled as properties of the substances, not reactions, and are
quoted as G
0
f
of compounds from their elements at 25°C and in their “standard states”
which are generally pure solids, pure liquids or pure gases at one atmosphere partial pres-
sure. Because biological reactions occur almost exclusively in aqueous solution, values for
DG
0
f
are also quoted at a standard concentration of 1 M in aqueous solution. Values for
some compounds of biological interest found in anaerobic treatment are listed in Table 1
(Mosey, 1985).
As mentioned before, using the data of Table 1 it is possible to estimate DG
0
of the half-
reactions. The standard redox potential (E
0
) is calculated from DG
0
of the half reaction as
E
0
= (DG
0
)/(n.F), where n is the number of electrons in the half-reaction and F is the
Faraday constant expressed as mV-equivalents (96.42 J/mV). In order to evaluate the redox
potential the convention is to write the half-reaction with the electrons on the left, as a
Water Science and Technology Vol 44 No 8 pp 77–84 © IWA Publishing 2001
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