23 © Springer Nature Switzerland AG 2019
R. Naraian (ed.), Mycodegradation of Lignocelluloses, Fungal Biology,
https://doi.org/10.1007/978-3-030-23834-6_2
Chapter 2
Fungal Biodiversity Producing Cellulase
Involved in Effcient Cellulolysis
Praveen Kumar Gupta, Shreya Choudhary, C. Chandrananthi,
J. Sharon Eveline, and S. P. Sushmitha
1 Introduction
Cellulases are a class of lignocellulolytic enzymes involved in converting cellulose
into simple sugars. Lignocellulose constitutes 60% of the plant cell wall and is one
of its major components. Lignocellulose comprises a family of lignin, hemicellu-
loses, and cellulose which are sugar biopolymer and its derivative (Galbe and Zacchi
2012). Cellulase is a polymer chain of D-glucose as monomeric units. It is the most
widely available organic polymer in the world having 4 × 10
10
tonnes produced
every year (Coughlan 1985). Cellulases from microbial sources have become the
main focus due to varied biotechnological applications. These are synthesized on
cellulosic material by various fungal and bacterial sources (aerobic, anaerobic,
mesophilic, or thermophilic) during their growth. Fungal cellulases are known to be
simpler than their bacterial counterpart having two separate domains. The two sepa-
rate domains are (1) catalytic domain and (2) cellulose-binding module (CBM),
having approximately 35 amino acids (Carvalho et al. 2003).
Cellulase is a 3-enzyme membered family each having a particular function. The
members in this family are exo-(1,4)-β-d-glucanase which is responsible for attack-
ing cellulose chain ends, endo-(1,4)-β-d-glucanase which acts on O-glycosidic
bonds in a random order which gives different lengths of glucan chain. The third is
β-glucosidases which produces cellulose by acting on β-cellobiose (Kuhad et al.
2011). The most widely used and studied microorganisms for cellulase production
are Trichoderma, Aspergillus, Cellulomonas, Penicillium, Clostridium, etc. (Kuhad
et al. 2011).
P. K. Gupta (*) · S. Choudhary · C. Chandrananthi · J. Sharon Eveline · S. P. Sushmitha
Department of Biotechnology, R.V College of Engineering, Bangalore, India
e-mail: praveenkgupta@rvce.edu.in